Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
From their detailed study of the flowering plants botanists have leaned
toward different groups as the present representatives of the primitive
types. The various claims of the different families to this position
cannot be considered here; probably that of the Ranales (the group of
families round Ranunculaceæ as a central type) is the best supported. Yet
these plants are most frequently delicate herbs, which would have stood
relatively less chance of fossilization than the other families which may
be considered primitive. They are peculiarly remote from the group of
Bennettiteæ in their vegetative structure, a fact the importance of which
seems to have been underrated, for in the same breath we are assured that
the Bennettites are a kind of cousin to the ancient Angiosperms, and that
the Ranales are among the most primitive living Angiosperms, and
therefore presumably nearest the ancient ones.
However, let us leave the charms of controversy on one side and look at
the actual structure of the group. They were widely spread in Lower
Mesozoic times, the plants being preserved as casts, impressions, and
with structure in great numbers. The bulk of the described structural
specimens have been obtained from the rocks of England, France, Italy,
and America, although leaf impressions are almost universally known. The
genus _Williamsonia_ belongs to this family, and is one of the best known
of Mesozoic plant impressions.
Externally the Bennettiteæ were identical in appearance with stumpy
Cycads, and their leaves it is which gave rise to the surmise, so long
prevalent, that the Lower Mesozoic was the “Age of Cycads”, just as it
was the Pteridosperm leaves that gave the Palæozoic the credit of being
the “Age of Ferns”. In the anatomy of both stem and leaf, also, the
characters are entirely Cycadean; the outgoing leaf trace is indeed
simpler in its course than that of the Cycads.
[Illustration: Fig. 71.—Half of a Longitudinal Section through a Mature
Cone of _Bennettites_
A, Short conical axis; _s_, enclosing bracts; S, seeds; _sc_, sterile
scales between the seeds.]
The fructifications, however, differ fundamentally from those of the
Cycads, as indeed they do from those of any known family. They took the
form of compact cones, which occurred in very large numbers in the mature
plants hidden by the leaf bases. In _Williamsonia_, of which we know much
less detail, the fructifications stood away from the main axis on long
pedicels.
Public-domain text, read in full here on John Shaqi.
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