Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
The best known of the genera is that of _Cycas_, of which an illustration
is given in fig. 74. The thick, stumpy stem and crown of “palm”-like
leaves give it a very different appearance from any other Gymnosperm.
Commonly the plants reach only a few feet in height, but very old
specimens may grow to the height of 30 ft. or more. The other genera are
smaller, and some have short stems and a very fern-like appearance, as,
for example, the genus _Stangeria_, which was supposed to be a fern when
it was first discovered and before fruiting specimens had been seen.
The large compound leaves are all borne directly on the main stem,
generally in a single rosette at its apex, and as they die off they leave
their fleshy leaf bases, which cover the stem and remain for an almost
indefinite number of years.
The wood of the main trunks differs from that of the other Gymnosperms in
being very loosely built, with a large pith and much soft tissue between
the radiating bands of wood. There is a cambium which adds zones of
secondary tissue, but it does not do its work regularly, and the cross
section of an old Cycad stem shows disconnected rings of wood,
accompanied by much soft tissue. The cells of the wood have bordered pits
on their walls, and in the main axis the wood is usually all developed in
a centrifugal direction, but in the axis of the cones some centripetal
wood is found (refer to _c_, fig. 65, p. 97).
[Illustration: Fig. 74.—Plant of _Cycas_, showing the main stem with
the crown of leaves and the irregular branches which come on an old
plant]
In their fructifications the Cycads stand even further apart from the
rest of the Gymnosperms. One striking point is the enormous size of their
_male_ cones. The male cones consist of a stout axis, round which are
spiral series of closely packed simple scales covered with pollen-bearing
sacs (which bear no inconsiderable likeness to fern sporangia), the whole
cone reaching 1½ ft. in length in some genera, and weighing several
pounds. All the other Gymnosperms, except the Araucareæ, where they are
an inch or two long, have male cones but a fraction of an inch in length.
[Illustration: Fig. 75.—Seed-bearing Scale of _Cycas_, showing its
lobed and leaflike character
_s_, Seeds attached on either side below the divisions of the
sporophyll.]
In all the members of the family, excepting _Cycas_ itself, the female
fructifications also consist of similarly organized cones bearing a
couple of seeds on each scale instead of the numerous pollen sacs. In
_Cycas_ the male cones are like those of the other genera, and reach an
enormous size; but there are no female cones, for the seeds are borne on
special leaflike scales. These are illustrated in fig. 75, which shows
also that there are not two seeds (as in the other genera with cones) to
each scale, but an indefinite number.
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