Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
First in the Permian we meet with the important genus of plant called
_Calamites_, which were very abundant in the Coal Measures. Many of the
Calamites were of great size, for specimens with large trunks have been
found 30 ft. and more long, which when growing must certainly have been
much taller than that. The number of individuals must also have been very
great, for casts and impressions of the genus are among the commonest
fossils. They were, in fact, one of the dominant groups of the period.
Like the Lycopods, the Equisetaceæ reached their high-water mark of
development in the Carboniferous period; at that time the plants were
most numerous, and of the largest size and most complicated structure
that they ever attained.
[Illustration: Fig. 104.—Small Branches attached to stouter Axis of
_Calamites_. Photo of Impression]
As will be immediately suspected from analogy with the Lycopods, they
differed from the modern members of the family in their strongly
developed anatomy, and in the strength and quantity of their secondary
wood. Yet in their external appearance they probably resembled the living
genus in all essentials, and the groves of the larger ones of to-day
growing in the marshes probably have the appearance that the palæozoic
plants would have had if looked at through a reversed opera glass.
Fig. 104 is a photograph of some of the small branches of a Calamite, in
which the ribbed stem can be seen, and on the small side twigs the fine,
pointed leaves lying in whorls.
In most of the fossil specimens, however, particularly the larger ones,
the ribs are not those of the true surface, but are those marked on the
_internal cast_ of the pith.
[Illustration: Fig. 105.—Transverse Section of _Calamites_ Stem with
Secondary Wood _w_ formed in Regular Radial Rows in a Solid Ring
_c_, Canals associated with the primary bundles; _p_, cells of the
pith, which is hollow with a cavity _l_, _cor_, Cortex and outer
tissues well preserved. (Microphoto.)]
Among tissue petrifactions there are many Calamite stems of various
stages of growth. In the very young ones there are only primary bundles,
and these little stems are like those of a living Equisetum in their
anatomy, and have a hollow pith and small vascular bundles with canals
associated. The fossil forms, however, soon began to grow secondary wood,
which developed in regular radial rows from a cambium behind the primary
bundles and joined to a complete ring.
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