Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
We have a general knowledge of the external appearance of _Sphenophyllum_
from the numerous impressions of leaves attached to twigs which are found
in the rocks of the Carboniferous period. These impressions present a
good deal of variety, but all have rather delicate stems with whorls of
leaves attached at regular intervals. The specimens are generally easy to
recognize from the shape of the leaves, which are like broad wedges
attached at the point (see fig. 112). In some cases the leaves are more
finely divided and less fanlike, and it may even happen that on the same
branch some may be wedge-shaped like those in fig. 112, and others almost
hairlike. This naturally suggests comparison with water plants, which
have finely divided submerged leaves and expanded aerial ones. In the
case of _Sphenophyllum_, however, the divided leaves sometimes come at
the upper ends of the stems, quite near the cones, and so can hardly have
been those of a submerged part. The very delicate stems and some points
in their internal anatomy suggest that the plant was a trailing creeper
which supported itself on the stouter stems of other plants.
[Illustration: Fig. 112.—Impression of _Sphenophyllum_ Leaves attached
to the Stem, showing the wedge-shaped leaflets arranged in whorls]
The stems were ribbed, but unlike those of the Calamites the ribs ran
straight down the stem through the nodes, and did not alternate there, so
that the bundles at the node did not branch and fuse as they did in
_Calamites_.
The external appearance of the long slender cones was not unlike that of
the Calamite cones, though their internal details showed important
distinctions.
In one noticeable external feature the plants differed from those of the
last two groups considered, and that was in their _size_. Palæozoic
Lycopods and Equisetaceæ reached the dimensions of great trees, but
hitherto no treelike form of _Sphenophyllum_ has been discovered, and in
the structure-petrifactions the largest stems we know were less than an
inch in diameter.
In the internal anatomy of these stems lies one of the chief interests
and peculiarities of the plants. In the very young stage there was a
sharply pointed solid triangle of wood in the centre (fig. 113), at each
of the corners of which was a group of small cells, the protoxylems. The
structure of such a stem is like that of a root, in which the primary
wood all grows inwards from the protoxylems towards the centre, and had
we had nothing but these isolated young stems it would have been
impossible to recognize their true nature.
[Illustration: Fig. 113.—_Sphenophyllum_, Transverse Section of Young
Stem
_c_, Cortex, the soft tissue within which has decayed and left a space,
in which lies the solid triangle of wood, with the small protoxylem
groups _px_ at each corner. (Microphoto.)]
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