Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
In its anatomy the stalk of the cone has certain features similar to
those in the stem proper, which were among the first indications that led
to the discovery that the cone belonged to _Sphenophyllum_. There were
numerous spores in each of the sporangia, which had coats ornamented with
little spines when they were ripe (fig. 117, if examined with a
magnifying glass, will show this). Hitherto the only spores known are of
uniform size, and there is no evidence that there was any differentiation
into small (male) and large (female) spores such as were found in some of
the Lepidodendrons. In this respect _Sphenophyllum_ was less specialized
than either _Lepidodendron_ or _Calamites_.
In the actual sections of _Sphenophyllum_ cones the numerous sporangia
seem massed together in confusion, but usually some are cut so as to show
the attachment of the stalk, as in fig. 117, _st_. As the stalk was long
and slender, but a short length of it is usually cut through in any one
section, and to realize their mode of attachment to the axis (as shown in
fig. 116) it is necessary to study a series of sections.
[Illustration: Fig. 117.—Part of Cone of _Sphenophyllum_, showing
sporangia _sp_, some of which are cut so as to show a part of their
stalks _st_. B, Bract. (Microphoto.)]
Of the other plants belonging to the group, _Bowmanites Römeri_ is
specially interesting. Its sporangia were borne on stalks similar to
those of _Sphenophyllum_, but each stalk had two sporangia attached to
it. Two sporangia are also borne on each stalk in _S. fertile_. These
plants help in elucidating the nature of the stalked sporangia of
_Sphenophyllum_, for they seem to indicate a direct comparison between
them and the sporophylls of the Equisetales.
There is, further, another plant, of which we only know the cone, of
still greater importance. This cone (_Cheirostrobus_) is, however, so
complex that it would take far too much space to describe it in detail.
Even a diagram of its arrangements is extraordinarily elaborate. To the
specialist the cone is peculiarly fascinating, for its very complexity
gives him great scope for weaving theories about it; but for our purposes
most of these are too abstruse.
[Illustration: Fig. 118.—A, Diagram of Three-lobed Bract from Cone of
_Cheirostrobus_. _a_, Axis; _br_, the three sterile lower lobes of the
bract; _sp_, the three upper sporophyll-like lobes, to each of which
were attached four sporangia S. B, Part of the above seen in section
longitudinal to the axis. (Modified from Scott.)]
Public-domain text, read in full here on John Shaqi.
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