Nor, again, is the complexity of the adjustment distinctive of
intelligence as contrasted with instinct. We have cited examples
which afford evidence of much complexity in instinctive behaviour.
The construction and storage of the nest among solitary wasps, and
their methods of capturing and conveying the insects or spiders on
which they prey, are sufficiently complex. So, too, is the behaviour
of the Sitaris larva which attaches itself to the male bee, passes to
the female, and then slips on to the eggs she lays; and so, again,
is that of the Yucca moth, which collects pollen from the anthers,
conveys it to the stigma, and then lays her eggs among the ovules.
These cases show, too, that the circumstances may be dealt with in
such a way as to extract from them the maximum of benefit. It would be
difficult intelligently to improve upon the manner of dealing with the
circumstances displayed in many familiar modes of instinctive procedure.
There remain the novelty of the adjustment and the individuality
displayed. And here we seem to have valid criteria of intelligent
behaviour. The ability to perform acts in special adaptation to
new circumstances, and the individuality manifested in dealing with
the complex conditions of a variable environment,--these seem to be
distinctive features of intelligence. On the other hand, in instinctive
behaviour there seems to be no choice; the animal is impelled to their
stereotyped performance through impulse, as by a stern necessity; they
are so far from novel that they are performed by every like individual
of the species, and have been so performed by their ancestors for
generations; and in performing the instinctive act, the animal seems to
have no more individuality or originality than a piece of adequately
wound clockwork.
Granting, then, that behaviour is shown to be intelligent by the fact
that there is evidence of profiting by experience, we may say that
the level attained by the intelligence is indicated by the complexity
of the adjustment, its precision, the individuality shown, the amount
of prevision disclosed, and in its being such as to extract from the
circumstances the maximum of benefit. Many of these points, however,
serve equally well to mark the level of instinctive procedure.
II.--INTELLIGENT BEHAVIOUR IN INSECTS
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