Here we have intelligent behaviour rising to a level to which some
would apply the term rational. For the act may be held to afford
evidence of the perception of the relation of the means employed to
an end to be attained, and some general conception of purpose. In
this section, which deals with description of behaviour based on
observation, the psychological explanation cannot be discussed. Similar
indications of deliberate action may be held to be afforded by the
sometimes elaborate “locality studies” which these insects seem to
make,--by the “care that is taken by wasps to acquaint themselves with
the surroundings of their nests.” A _Sphex_, for example, which had
partially made and then abandoned several nests, left them without any
locality study; but when she had completed a nest in a suitable spot
she made “a most thorough and systematic study of the surroundings.
She flew in and out among the plants, first in narrow circles near
the surface of the ground, and then in wider and wider ones as she
rose higher in the air, until at last she took a straight line and
disappeared in the distance.” Another species (_Cerceris deserta_) “has
the habit of making a number of half circles in front of the nest, and
then, after rising a little higher, of flying several times completely
round it.” The method of procedure is, it seems, so normal to the
species that it is probably founded on an instinctive basis. Dr. and
Mrs. Peckham, in commenting on their observations, say:[54] “If the
examination of the objects about the nest makes no impression upon the
wasp, or if it is not remembered, she ought not to be inconvenienced
nor thrown off her track when weeds and stones are removed and
the surface of the ground is smoothed over; but this is just what
happens.” For convenience of observation they “sometimes gently moved
intercepting objects to one side, but even such a slight change threw
the wasp out of her bearings, and made it difficult for her to recover
her treasure.” Where wasps form a number of nests in a small plot of
ground, as in the case of _Bembex_, each knows and returns to its own
hole, as was proved by Dr. Peckham, who marked the insects and their
nests with paint.
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