Courtship may thus be regarded from the physiological point of view
as a means of producing the requisite amount of pairing-hunger; of
stimulating the whole system and facilitating general and special
vascular changes; of creating that state of profound and explosive
instability which has for its psychological concomitant or antecedent
an imperious and irresistible craving. This not only overcomes the
coyness of the female, but generates and strengthens the ardour of
the male--a point on which, perhaps, Professor Groos does not lay
sufficient stress. For the process is reciprocal; and though the male
leads in the ardour of courtship, yet this ardour constantly grows till
at last it overcomes the barriers of reluctance. Courtship is thus
the strong and steady bending of the bow that the arrow may find its
mark in a biological end of the utmost importance in the survival of a
healthy and vigorous race.
The coyness and reluctance of the female afford the conditions under
which the bow is bent to the full. But they also afford the conditions
of the apparent act of choice. This takes the place, on the perceptual
plane of mental development, of that deliberation which precedes the
higher act of choice on the ideational plane. For psychology, as well
as for biology, then, Dr. Groos’s suggestion is a welcome and helpful
one. Both upholders of sexual selection and critics of that hypothesis,
have been apt to regard the choice of a mate in animals from too
anthropomorphic a point of view--to look upon it as the outcome of
rational deliberation, of weighing in the æsthetic balance the relative
attractiveness of this suitor and of that, and of reaching a definite
conclusion that the one is to be accepted because he behaves or is
adorned in such and such a way, while the other is to be rejected
because he falls below all reasonable standards of requirement. The
choice exercised by the female, if so we term it, is far simpler and
more naïve. Indeed, Professor Groos goes so far as to say that on his
hypothesis the element of choice is altogether abolished. “It is the
instinctive coyness of the female,” he says,[152] “that necessitates
all the arts of courtship, and the probability is that seldom or never
does the female exert any choice. She is not awarder of the prize, but
rather a hunted creature. So, just as the beast of prey has special
instincts for finding his prey, the ardent male must have special
instincts for subduing female reluctance. According to this theory,
there is choice only in the sense that the hare finally succumbs to the
best hound, which is as much as to say that the phenomena of courtship
are referred at once to natural selection.” He reaches this conclusion,
however, by gradual stages. He first urges[153] that “it would be
absurd to affirm that all bird songs originate in a conscious æsthetic
and critical act of judgment on the part of the female. A conscious
choice either of the most beautiful or the loudest singer is certainly
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