Leaving on one side, then, the natural origin of protoplasm, the
conditions of which are unknown, we find that, as a matter of
observation, every bit of living substance, the history of which has
been traced, is a fragment detached from some other bit which behaved
in the same way. This is the basal fact of the continuity of organic
evolution. But such a detached fragment has the property of increasing
by taking up from the environment more of those elementary materials
from which it is itself compounded in subtle synthesis. Nay, further,
every fragment of which we know the history is found to increase in
such a way as to reach, in form, structure, and idiosyncracies of
behaviour, the likeness of the organism--plant or animal--from which it
was derived. In the higher plants and animals the separated fragments
or cells are the ova and sperms, or their equivalents, which unite,
with fusion or coalescence of their nuclear matter, and thus give rise
to a new individual in the course of embryological development.
Now, as we have already seen, much modern biological discussion centres
round the question whether the detached reproductive fragment, ovum or
sperm as the case may be, is derived from the whole body of the parent,
by what Darwin termed pangenesis or in some other way, or only from
germinal substance set apart in development for this end. And we have
provisionally accepted the hypothesis that it is the direct descendant
of other reproductive cells; and that, throughout a long ancestry,
stretching back into the far past, there never occurs in the direct
line of genealogical sequence, any highly differentiated cell, such as
a gland-cell, muscle-cell, nerve-cell; never, with certain reservations
into which we need not enter, is found the representative of any
tissue save that to which the reproductive function is restricted. In
technical phraseology, the continuity of organic evolution is due to
the continuity of the germinal substance.
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