Even in the case of the very first exhibition of such a deferred
instinct as the moor-hen’s dive, although that organized sequence
of acts which constituted the behaviour as a whole had never before
occurred, although there was no gradual learning how to dip beneath
the surface, and to swim under water, still many of the constituent
acts had been often repeated; experience had already been gained of
much of the detail then for the first time combined in an instinctive
sequence. So that if we distinguish between instinct as congenital and
habit as acquired, we must not lose sight of the fact that there is
continual interaction, in a great number of cases, between instinct
and habit, and that the first performance of a deferred instinct may
be carried out in close and inextricable association with the habits
which, at the period of life in question, have already been acquired.
Instinct supplies an outline sketch of behaviour, to which experience
adds colour and shading. Which predominates in the finished picture
depends on the status of the animal. In the lower and less intelligent
types the outline stands out clearly, there being but little shading to
divert our attention from the clear firm lines inscribed by heredity;
but in the higher and more intelligent animals, the deft pencil of
experience has added so much detail and has interwoven with the fainter
outline so many new and skilfully introduced touches, that the original
sketch is scarcely distinguishable unless we have carefully watched
from the beginning the gradual development of the picture.
V.--THE EVOLUTION OF INSTINCTIVE BEHAVIOUR
It may be assumed that the fact of evolution is generally admitted.
The question of its method is, however, still open to discussion. It
is possible that, as some biologists contend, there is an inherent
tendency in organic beings to evolve in certain definite directions
independently of their relation to the environment. But it is scarcely
probable that instinctive behaviour is mainly due to any such inherent
tendency--of the nature of which in any case we know but little.
Setting this on one side, therefore, we have two hypotheses: first,
that instincts are the result of natural selection; secondly, that they
are due to the inheritance of acquired habits. These two views we will
now proceed to consider.
Public-domain text, read in full here on John Shaqi.
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