_Every response or change in response of an animal is then the result of the interaction of its original knowable nature and the environment._ This may seem too self-evident a corollary for mention. It should be so, but, unfortunately, it is not. Two popular psychological doctrines exist in defiance of it. One is the doctrine that the movements of early infancy are random, the original nature of the animal being entirely indifferent as to what movement shall be made upon a given stimulus. But no animal can have an original nature that does not absolutely prescribe just what the response shall be to every stimulus. If the movements are really random, they occur by virtue of some force that works at random. If the movements are really the result of the action of the environment on the animal’s nature, they are never random. A baby twiddles his thumbs or waves his legs for exactly the same sort of reason that a chick pecks at a worm or preens its wing. The other doctrine which witnesses to neglect of the axiom that behavior is the creation of the environment, acting on the animal’s nature, is the doctrine that the need for a certain behavior helps to create it, that being in a difficulty tends in and of itself to make an animal respond so as to end the difficulty. The truth is that to a difficulty the animal responds by whatever its inherited and acquired nature has connected with the special form of difficulty and that in many animals the one response of those thus provided which relieves the difficulty is selected and connected more firmly with that difficulty’s next appearance. The difficulty acts only as a stimulus to the animal’s nature and its relief acts only as a premium to the connection whereby it was relieved. The law of original behavior, or the law of instinct, is then that _to any situation an animal will, apart from learning, respond by virtue of the inherited nature of its reception-, connection- and action-systems_. The inquiry into the laws of learning to be made in this essay is limited to those aspects of behavior which the term has come historically to signify, that is, to intellect, skill, morals and the like. For the purposes of this essay it is not necessary to decide just what features of an animal’s behavior to include under intellect, skill, morals and the like. The statements to be made will fit any reasonable dividing line between behavior on the one side and mere circulation, digestion, excretion and the like on the other. There should in fact be no clear dividing line, since there is no clear gap between those activities which naturalists have come to call behavior and the others.