So far as heredity is concerned, it matters not whether there be a
continuity of germ-cells or of germ-plasma. In either case, the
essential feature is that body-cells as such cannot give rise to the
germ--that the hen cannot produce the egg. On either view, characters
acquired by the body cannot be transmitted to the offspring through the
ova or sperms. The annexed diagram illustrates how, on the view that the
hen produces the egg, dints hammered into the body by the environment
will be handed on; while, on the view that the hen does not produce the
egg, the dints of the environment are not transmitted to the offspring.
On the hypothesis of continuity, heredity is due to the fact that two
similar things under similar conditions will give similar products. The
ovum from which the mother is developed, and the ovum from which the
daughter is developed, are simply two fragments separated at different
times from the same continuous germ-plasm.[BO] Both develop under
similar circumstances, and their products cannot, therefore, fail to be
similar. How variation is possible under these conditions we shall have
to consider presently.
[Illustration: Fig. 21.--Egg and hen.
I. "The egg produces the hen." II. "The hen produces the egg." In I. the
dints produced by the environment are not inherited; in II. they are.
The letters indicate successive individuals. The small round circles
indicate the eggs.]
Now, although I value highly Professor Weismann's luminous researches,
and read with interest his ingenious speculations, I cannot but regard
his doctrine of the continuity of germ-plasm as a distinctly retrograde
step. His germ-plasm is an unknowable, invisible, hypothetical entity.
Material though it be, it is of no more practical value than a
mysterious and mythical germinal principle. By a little skilful
manipulation, it may be made to account for anything and everything. The
fundamental assumption that whereas germ-plasm can give rise to
body-plasm to any extent, body-plasm can under no circumstances give
rise to germ-plasm, introduces an unnecessary mystery. Biological
science should set its face against such mysteries. The fiction of two
protoplasms, distinct and yet commingled, is, in my opinion, little
calculated to advance our knowledge and comprehension of organic
processes. For myself, I prefer to take my stand on protoplasmic unity
and cellular continuity.
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