(3) It is sometimes said, in bold metaphor, that natural selection is
constantly on the watch to select any modification, however slight,
which is of advantage to the species. And it is true that elimination is
ceaselessly operative. But it is equally certain that the advantage must
be of sufficient value to decide the question whether its possessor
should be eliminated or should escape elimination. If it does not reach
this value, Natural Selection, watch she never so carefully, can make no
use of it. Elimination need not, however, be to the death; exclusion
from any share in continuing the species is sufficient. To breed or not
to breed, that is the question. Any advantage affecting this essential
life-function will at once catch the eye of a vigilant natural
selection. But it must be of sufficient magnitude for the machinery of
natural selection to deal with. That machinery is the elimination of a
certain proportion of the individuals which are born. Which shall be
eliminated, and which shall survive, depends entirely on the way in
which the individuals themselves come out in life's competitive
examination. The manner in which that examination is conducted is often
rude and coarse, too rough-and-ready to weigh minute and infinitesimal
advantages.
What must be the value of a favourable or advantageous modification to
decide the question of elimination, to make it an _available advantage_,
must remain a matter of conjecture. It will vary with the nature and the
pressure of the eliminative process. And perhaps it is scarcely too much
to say that, at present, we have not observational grounds on which to
base a reliable estimate in a single instance. We must not let our
conviction of its truth and justice blind us to the fact that natural
selection is a logical inference rather than a matter of direct
observation. A hundred are born, and two survive; the ninety-eight are
eliminated in the struggle for existence; we may therefore infer that
the two escaped elimination in virtue of their possession of certain
advantageous characters. There is no flaw in the logic that has thus
convinced the world that natural selection is a factor in evolution. But
by what percentage of elimination-marks the second of the two successful
candidates beats the senior on the list of failures we do not know. We
can only see that, on the hypothesis of natural selection, it must have
been sufficiently appreciable to determine success or failure.
Public-domain text, read in full here on John Shaqi.
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