Under natural selection, as we have seen, the development of colour is
fostered under certain conditions. The colour is either protective,
rendering the organism inconspicuous amid its normal surroundings, or it
is of warning value, advertising the organism as inedible or dangerous,
or, in the form of recognition-marks, it is of service in enabling the
members of a species to recognize each other. Now, in the case of both
warning colour and recognition-marks, their efficacy depends upon the
perceptual powers of animals. Unless there be a rapidly acquired and
close association of the quality we call nastiness with the quality we
call gaudiness (though, for the animal, there is no such _isolation_ of
these qualities as is implied in our words [CV]), such that the sight of
the gaudy insect suggests that it will be unpleasant to eat, the
gaudiness will be of no avail. And if there is any truth in the doctrine
of mimicry, the association is particular. It is not merely that bright
colours are suggestive of a nasty taste. The insect-eating birds
associate nastiness especially with certain markings and
coloration--"the tawny _Danais_, the barred _Heliconias_, the blue-black
_Euplæas_, and the fibrous _Acræas_;" and this is proved by the fact
that sweet insects mimicking these particular forms are thereby
protected.
So, too, with recognition-marks. If the bird or the mammal have not
sufficient perceptive powers to distinguish between the often not very
different recognition-marks, of what service can they be?
Recognition-marks and mimicry seem, therefore, to show that in the
former case many animals, and in the latter the insect-eating birds,
mammals, lizards, and other animals concerned, have considerable powers
of perception and association.
Among other associations are those which are at the base of what I have
termed preferential mating. We must remember how deeply ingrained in the
animal nature is the mating instinct. _We_ may find it difficult to
distinguish closely allied species. But the individuals of that species
are led to mate together by an impelling instinct that is so well known
as to elicit no surprise. Instinct though it be, however, the mating
individuals must recognize each other in some way. The impulse that
draws them together must act through perceptual agency. It is not
surprising, therefore, to find, when we come to the higher animals,
that, built upon this basis, there are well-marked mating preferences.
And this, as we have before pointed out, following Wallace, is an
efficient factor in segregation. Let us, however, hear Mr. Wallace
himself in the matter.
Public-domain text, read in full here on John Shaqi.
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