6. Variations in the time at which reproduction commences.
7. Variations in the duration and amount of parental protection and
fosterage.
8. Variations in the period at which secondary sexual characters and the
maximum efficiency of the several organs is reached.
It is impossible here to discuss these modes of variation _seriatim_. I
shall therefore content myself with but a few remarks on the importance
of protection and fosterage. It is not too much to say that, without
fosterage and protection, the higher forms of evolution would be
impossible. If you are to have a highly evolved form, you must allow
time for its evolution from the egg; and that development may go on
without let or hindrance, you must supply the organism with food and
lighten the labour of self-defence. Most of the higher organisms are
slow in coming to maturity, passing through stages when they are
helpless and, if left to themselves, would inevitably fall a prey to
enemies.
In those animals in which the system of fosterage and protection has not
been developed a great number of fertilized ova are produced, only a few
of which come to maturity. It might be suggested that this is surely an
advantage, since the greater the number produced the greater the chances
of favourable variations taking place. But it has before been pointed
out that these great numbers are decimated, and more than decimated, not
by elimination, but by indiscriminate destruction; embryos, good, bad,
and indifferent, being alike gobbled up by those who had learnt the
secret of fostering their young. The alternative has been between
producing great numbers[DN] of embryos which soon fend for themselves,
and a few young who are adequately provided for during development. And
the latter have proved the winners in life's race. If we compare two
flat-fishes belonging to very different groups, the contrast here
indicated will be readily seen. The skate is a member of the shark
tribe, flattened symmetrically from above downwards. It lays, perhaps,
eighty to a hundred eggs. Each of these is large, and has a rich supply
of nutritive food-yolk. Each is also protected by a horny case with
pointed corners--the so-called sea-purse of seaside visitors. These are
committed by the skate to the deep, and are not further cared for. But
the abundant supply of food-yolk gives the little skate which emerges a
good start in life. On the other hand, the turbot, one of the bony
fishes, flattened from side to side with an asymmetrical head, lays
several millions of eggs, which float freely in the open sea. These are
minute and glassy, and not more than one-thirtieth of an inch in
diameter. When the fishes are hatched, they are not more than about
one-fifth of an inch in length. The slender stock of food-yolk is soon
used up, and henceforth the little turbot (at present more like a
stump-nosed eel than a turbot) has to get its own living. Hundreds of
thousands of them are eaten by other fishes.
Public-domain text, read in full here on John Shaqi.
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