The point, I repeat, to be especially noted about these cases and others
which might be cited,[EM] is that the variation produced is a definite
variation. Very probably it is generally, or perhaps always, produced in
the embryonic or larval period of life. In some cases the variation
seems to be transmissible, though definite and satisfactory proofs of
this are certainly wanting. Still, we may say that if the changed
conditions be maintained, the resulting variation will also be
maintained. Under these conditions, at least, the variation is a stable
one. It is probable that, apart from preferential mating, the varieties
thus produced will tend to breed together rather than to be crossed with
the parent form or varieties living under different conditions. In this
way varieties may sometimes arise by definite and perhaps considerable
leaps under the influence of changed conditions. We must not run the
adage, _Natura nil facit per saltum_, too hard, nor interpret _saltum_
in too narrow a sense.
It is true, and we may repeat the statement of the fact for the sake of
emphasis, that we do not know how or why this or that particular
variation should result from this or that change of climate,
environment, or food-stuff; nor do we know why certain variations (such
as that which produced the ancon breed of sheep) should be stable, while
other variations are peculiarly unstable. But in this we are not worse
off than we are in the study of inorganic nature. We do not know why
calcite should crystallize in any particular one of its numerous
varieties of crystalline form; we do not know why some of these are more
stable than others. We may be able to point to some of the conditions,
but we cannot be said to understand why arragonite should be produced
under some circumstances, calcite under others; or why the same
constituents should assume the form of augite in some rocks, and
hornblende in other rocks. We are hedged in by ignorance; and perhaps
one of our chief dangers, becoming with some people a besetting sin, is
that of pretending to know more than we are at present in a position to
know. Our very analogies by which we endeavour to make clear our meaning
may often seem to imply an unwarrantable assumption of knowledge.
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