Whether sexual selection is also a factor is still a matter of opinion.
Without it the specific character and constancy of secondary sexual
features are at present unexplained. If inherited use and disuse are
admitted as factors in origin, they must also be admitted as important
factors in guidance.
Questions of origin and guidance should, so far as is possible, be kept
distinct. These terms, however, apply to the origin and guidance of
variations. In the origin of species guidance is a factor, no doubt a
most important factor. The title of Darwin's great work was, therefore,
perfectly legitimate. And those who say that natural selection plays no
part in the origin of species are, therefore, undoubtedly in error.
NOTES
[CI] It is beyond the scope of this book to give the _evidences_ of
evolution. Such evidence from embryology, from distribution, and
from palæontology, is now abundant. For palæontological evidence,
see Nicholson's "Manual of Palæontology," 3rd edit., especially
the second volume on "Vertebrates," by R. Lydekker.
[CJ] Weismann, "Essays on Heredity," p. 24.
[CK] Ibid. p. 140.
[CL] Weismann, "Essays on Heredity," p. 90.
[CM] Ibid. p. 292. See also a discussion in _Nature_, in which Mr.
Romanes and Professor Ray Lankester took part, beginning vol. xli.
p. 437.
[CN] Weismann, "Essay on Heredity," p. 140.
[CO] "Origin of Species," p. 110.
[CP] With regard to blind cave-fish, Professor Ray Lankester has
suggested that some selection has been effected. Those animals
whose sight-sensitiveness enabled them to detect a glimmer of
light would escape to the exterior, leaving those with
congenitally weak sight to remain and procreate in the darkness of
the cave.
[CQ] Darwin, "Descent of Man," pt. ii. chap. viii.
[CR] "Darwinism," chap. x.
[CS] "Darwinism," p. 295. Messrs. Geddes and Thomson, "The Evolution of
Sex," p. 28, also contend that "combative energy and sexual beauty
rise _pari passu_ with male katabolism."
[CT] "Darwinism," p. 293.
[CU] Mr. Poulton, who takes a similar line of argument in his "Colours
of Animals," lays special stress upon the production of _white_
(see p. 326).
[CV] See Chapter VIII.
[CW] "Darwinism," p. 172.
[CX] See "Animals and Plants under Domestication," vol. ii. p. 80.
[CY] "Darwinism," p. 332.
[CZ] "The Colour-Sense," by Grant Allen, p. 95.
[DA] That on "The Emotions of Animals" (X.).
[DB] "Darwinism," p. 318.
[DC] Natural History Society of Wisconsin, vol. i. (1889).
[DD] "Darwinism," p. 286.
[DE] On the negative character of disuse, see p. 196.
[DF] Cope, "Origin of the Fittest," p. 374.
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