The organ of vision, then, in us consists of an essential sensory
membrane, the retina, with its delicate rods and cones; and an accessory
apparatus for focussing an inverted image on to the sensitive surface of
the retina. The surface is not, however, equally sensitive, or, in any
case, does not give an equal power of discrimination, throughout its
whole extent. This is seen in the experiment above described. When we
look at the dot we see the coin, but not distinctly. The area of clear
and distinct vision is, in fact, very small, constituting the yellow
spot about 1/12 of an inch (2 millimetres) long, and 1/30 of an inch (.8
millimetre) broad. And even within this small area there is a still more
restricted area of most acute sensibility only 1/120 of an inch (.2
millimetre) in diameter. Nevertheless, within this minute area there are
some two thousand cones, the rods being here absent. In carefully
examining an object we allow this area of acute vision to range over it.
Hence the extreme value of that delicate mobility which the eye
possesses--a mobility that is accompanied by muscular sensations of
great nicety.
We saw that the sense of touch in the tongue is sufficiently delicate to
enable us to recognize, as two, points of contact separated by 1/25 of
an inch (1.1 millimetre). What, in similar terms, is the delicacy of
sight? At what distance apart, on the most delicate part of the retina,
can two points of stimulation be recognized as distinct from each other?
If the points of stimulation be not less than 1/6000 of an inch (.004
millimetre) apart, they can be distinguished as two. Below this they
fuse into one. The diameter of the end of a single cone in the yellow
spot is also about 1/6000 of an inch (.0045 millimetre).
With regard to the mode in which the stimulation of the retinal elements
is effected, we have no complete knowledge. Certain observations of Boll
and Kühne, however, show that when an animal is killed in the dark the
retina has a peculiar purple colour which is at once destroyed if the
retina be exposed to light. If a rabbit be killed at the moment when the
image, say, of a window, is formed on the retina, and the membrane at
once plunged in a solution of alum, the image may be fixed, and an
"optogram" of the window may be seen on the retina. The discharge of the
colour of the retinal purple may be regarded as the sign of a chemical
change effected by the impact of the light-vibrations. But in the yellow
spot there seems to be no visual purple. It is, indeed, developed only
in the rods, not in the cones. Here, probably, chemical or metabolic
changes occur without the obvious sign of the bleaching of retinal
purple. In the dusk-loving owl the retinal purple is well developed, but
in the bat it is said to be absent.
Public-domain text, read in full here on John Shaqi.
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