Unfortunately, all these are called indiscriminately eyes, or organs of
vision. An infusorian or a snail is said to see. But the terms "eye,"
"vision," "sight," imply that final excellence to which only the higher
animals, each on its own line, have attained.
This final excellence probably has its basis and earliest inception in
the fact that the functional activity of protoplasm is heightened in the
presence of ætherial vibrations. If, then, we imagine, as a
starting-point, a primitive transparent organism with a general
susceptibility to the influence of light-vibrations, the formation
within its tissues of pigment-granules absorbent of light will render
the spots where they occur specially sensitive to the ætherial
vibrations. Special refraction-globules would also act as minute lenses,
focussing the light, and thus concentrating it upon certain spots.
[Illustration: Fig. 39.--Direction-retina.
Simple retina for distinguishing the direction of the source of light or
of shadow.]
In many of the lower animals we find such organs, belonging to our first
category, and constituting either eye-spots of pigmented material or
simple lenses covering a pigmented area. If we call these eyes, we must
remember that in all probability they have no power of what we call
vision--only a power of distinguishing light from dark. Where, however,
there exists beneath the lens a so-called retina, that is, a layer of
rod-like endings of a nerve, it might, at first sight, be thought that
there, at any rate, we have true vision. But in all probability, in a
great number of cases the retinal rods are simply for the purpose of
rendering the organism sensitive, not only to the presence of light, but
to its direction. Light straight ahead (_a_) stimulates the middle rods;
from one side (_b_, _c_) it is focussed on the rods of the opposite side
of the retina; and similarly for intermediate positions. The presence of
a retinal layer is thus no infallible sign of a power of vision as apart
from mere sensibility to light. Indeed, in a great number of cases, from
the convexity and position of the lens, the formation of an image is
impossible. Only when it can be shown that a more or less definite image
can be focussed on the retina, or can be formed on the principle of
mosaic vision, can we justly surmise that a power of true vision is
present. I doubt whether this can be shown to be unquestionably the case
in any forms but the higher arthropods, the cuttle-fishes and their
allies, and the vertebrates.
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