[Illustration: Fig. 17.--Variations adjusted to the standard of the
noctule.]
Compared with this as a standard, the mean length of the second
metacarpal in the seven species is forty-three millimetres; that of the
third metacarpal, forty-four millimetres; and so on. The amount by which
each species exceeds or falls short of the mean is shown on the table,
and the points are joined up as before. Here, again, the table gives the
actual measurements in each case. For example, if the mean length of the
third metacarpal of the greater horseshoe bat be required, it is seen by
the table to fall short of the mean by four horizontal divisions and a
quarter, that is to say, by eight millimetres and a half. The length is
therefore (44 - 8-1/2) 35.5 millimetres.
Now, it will be seen from the table that the variation in the mean
length of the bones in different species is much greater than the
individual variations in the members of the same species. The table also
brings out in an interesting way the variation in the general character
of the wing. The noctule, for example, is especially strong in the
development of the second and third metacarpals, the phalanges of the
third digit being also a little above the average. Reference to the
figure of the bat's wing on p. 64 will show that these excellences give
length to the wing. It fails, however, in the metacarpal and phalanges
of the fifth digit, and in the length of the hind leg as represented by
the tibia. On consulting the figure of the wing, it is seen that these
are the bones which give breadth to the wing. Here the noctule fails.
Its wing is, therefore, long and narrow. It is a swallow among bats.
On the other hand, the horseshoe bats fail conspicuously in the second
and third metacarpals, though they make up somewhat in the corresponding
digits. On the whole, the wing is deficient in length. But the phalanges
of the fourth and fifth digits, and the length of the hind limb
represented by the tibia, give a corresponding increase of breadth. The
wing is, therefore, relatively short and broad. The long-eared bat,
again, has the third metacarpal and its digits somewhat above the mean,
and therefore a somewhat more than average length. But it has the fifth
metacarpal with its digit and also the tibia decidedly above the mean,
and therefore more than average breadth. Without possessing the great
length of the noctule's wing, or the great breadth of that of the
horseshoe, it still has a more than average length and breadth.
Public-domain text, read in full here on John Shaqi.
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