I venture to think that there is a more logical division than this. A
man who is dealing with animals or plants under domestication may
proceed by one of two well-contrasted methods. He may either select the
most satisfactory individuals or he may reject the most unsatisfactory.
We may term the former process _selection_, the latter _elimination_.
Suppose that a gardener is dealing with a bed of geraniums. He may
either pick out first the best, then the second best, then the third,
and so on, until he has selected as many as he wishes to preserve. Or,
on the other hand, he may weed out first the worst, then in succession
other unsatisfactory stocks, until, by eliminating the failures, he has
a residue of sufficiently satisfactory flowers. Now, I think it is clear
that, even if the ultimate result is the same (if, that is to say, he
selects the twenty best, or eliminates all but the twenty best), the
method of procedure is in the two cases different. Selection is applied
at one end of the scale, elimination at the other. There is a difference
in method in picking out the wheat-grains (like a sparrow) and
scattering the chaff by the wind.
Under nature both methods are operative, but in very different degrees.
Although the insect may select the brightest flowers, or the hen-bird
the gaudiest or most tuneful mate, the survival of the fittest under
nature is in the main the net result of the slow and gradual process of
the elimination of the unfit.[V] The best-adapted are not, save in
exceptional cases, selected; but the ill-adapted are weeded out and
eliminated. And this distinction seems to me of sufficient importance to
justify my suggestion that _natural selection_ be subdivided under two
heads--_natural elimination_, of widespread occurrence throughout the
animal world; and _selection proper_, involving the element of
individual or special choice.
The term "natural elimination" for the major factor serves definitely to
connect the natural process with that struggle for existence out of
which it arises. The struggle for existence is indeed the reaction of
the organic world called forth by the action of natural elimination.
Organisms are tending to increase in geometrical ratio. There is not
room or subsistence for the many born. The tendency is therefore held in
check by elimination, involving the struggle for existence. And the
factors of elimination are three: first, elimination through the action
of surrounding physical or climatic conditions, under which head we may
take such forms of disease as are not due to living agency; secondly,
elimination by enemies, including parasites and zymotic diseases; and
thirdly, elimination by competition. It will be convenient to give some
illustrative examples of each of these.
Public-domain text, read in full here on John Shaqi.
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