Sufficient has now been said concerning the modes of isolation and
segregation, geographical, preferential, and physiological. We must now
consider their effects. Where the isolated varieties are under different
conditions of life, there will be, through the elimination of the
ill-adapted in each case, differential adoption to these different
conditions. But suppose the conditions are similar: can there be
divergence in this case? The supposition is a highly hypothetical one,
because it postulates that all the conditions, climatal, environmental,
and competitive, are alike, which would seldom, if ever, be likely to
occur. Let us, however, make the supposition. Let us suppose that an
island is divided into two equal halves by the submersion of a stretch
of lowland running across it. Then the only possible causes of
divergence would lie in the organisms themselves[AW] thus divided into
two equal groups. We have seen that variations may be advantageous,
disadvantageous, or neutral. The neutral form a fluctuating, unfixed,
indefinite body. But they afford the material with which nature may
make, through intercrossing, endless experiments in new combinations,
some of which may be profitable. Such profitable variations would escape
elimination, and, if not bred out by intercrossing, would be preserved.
In any case, the variety would tend to advance through elimination as
previously indicated. But in the two equal groups we are supposing to
have become geographically isolated, the chances are many to one against
the same successful experiments in combination occurring in each of the
two groups. Hence it follows that the progress or advance in the two
groups, though analogous, would not be identical, and divergence would
thus be possible under practically similar conditions of life.
In his observations on the terrestrial molluscs of the Sandwich Islands,
Mr. Gulick notes that different forms are found in districts which
present essentially the same environment, and that there is no greater
divergence when the climatic conditions are dissimilar than there is
when those conditions are similar. As before noticed, the degree of
divergence is, roughly speaking, directly as the distance the varietal
forms are apart. Again, Darwin notes that the climate and environment in
the several islands of the Galapagos group are much the same, though
each island has a somewhat divergent fauna and flora. These facts lend
countenance to the view that divergence can and does occur under similar
conditions of life, if there be isolation. They seem, also, so far as
they go, to negative the view that the species is moulded directly by
the external conditions. For, if this factor were powerful, it would
override the effects of experimental combination of characters when the
conditions were similar, and would give rise to well-marked varietal
forms when the conditions were diverse.
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