If the body of a flying animal be in a horizontal position, a wing attached to it in such a manner that its under surface shall look forwards, and make an upward angle of 45° with the horizon is in a position to be applied either vertically (figs. 82 and 83, p. 158), or horizontally (figs. 67, 68, 69, and 70, p. 141). Such, moreover, is the conformation of the shoulder-joint in insects, bats, and birds, that the wing can be applied vertically, horizontally, or at any degree of obliquity without inconvenience.[118] It is in this way that an insect which may begin its flight by causing its wings to make figure-of-8 horizontal loops (fig. 71, p. 144), may gradually change the direction of the loops, and make them more and more oblique until they are nearly vertical (fig. 73, p. 144). In the beginning of such flight the insect is screwed _nearly vertically upwards_; in the middle of it, it is screwed _upwards and forwards_; whereas, towards the end of it, the insect advances in _a waved line_ almost horizontally (see _q´_, _r´_, _s´_, _t´_ of fig. 72, p. 144). The muscles of the wing are so arranged that they can propel it in a horizontal, vertical, or oblique direction. It is a matter of the utmost importance that the direction of the stroke and the nature of the angles made by the surface of the wing during its vibration with the horizon be distinctly understood; as it is on these that all flying creatures depend when they seek to elude the upward resistance of the air, and secure a maximum of elevating and propelling power with a minimum of slip. [118] The human wrist is so formed that if a wing be held in the hand at an upward angle of 45°, the hand can apply it to the air in a vertical or horizontal direction without difficulty. This arises from the power which the hand has of moving in an upward and downward direction, and from side to side with equal facility. The hand can also rotate on its long axis, so that it virtually represents all the movements of the wing at its root.