It is important to remark that platycnemy has been regarded as a
pithecoid structure, and for this reason the attempt has been made
to establish the degraded position of those peoples which are most
remarkable for platycnemy. But, as Boyd-Dawkins has already observed,
although the tibiæ of the gorilla and the chimpanzee are to some
extent platycnemic, they are much less so than the platycnemic bones
of the human skeleton. The tibia of a male gorilla in the College of
Surgeons Museum has an index width of 68·1, that of a female of 65·0,
while the index of the chimpanzee’s tibia is 61·1, which is about the
average of the tibias of Perthichwareu. It is unnecessary to indicate
the other marked distinctions between the tibiæ of men and apes; if
platycnemy is to be regarded as genetic, it must be admitted that man
has in this particular far exceeded apes.[57] Neither the gorilla,
the chimpanzee, the orang-utan, nor even the baboon possesses a tibia
which is flattened in its upper or middle part. In all these apes the
middle of the bone is more or less rounded, almost as if it had been
rounded by a turning-lathe. According to my experience, the degree of
platycnemy in anthropoids is subject to certain variations. It appears
to me to be least marked in the aged male gorilla (Fig. 41), and in
the gibbon (_Hylobates agilis_, _syndactylus_), in which latter animal
the transverse section of the tibia represents an almost equilateral
triangle. The platycnemy was more marked in an almost adult female
gorilla, still more decided in an aged male chimpanzee, which came
from the river Kiulu, and again in an aged female chimpanzee. On the
other hand, the centre of the shaft of the tibia in another aged male
chimpanzee which came from Loango, was rounded, and not platycnemic. In
the tibia of an adult orang-utan which I examined, the platycnemy was
very marked. But I agree with Boyd-Dawkins in never having met with an
anthropoid in which the platycnemy is so considerable as it is, for
instance, in the Cro-Magnon tibia, and in another found at Troy.
[Illustration:
Fig. 46.--Skeleton of the human foot, seen from above. _a_,
Astragalus. _b_, Os calcis. _c_, Scaphoid bone. _d_, _e_, _f_,
Cuneiform bones. _g_, Cuboid bone. _h_, Metatarsal bones. _ii_,
Phalanges.
]
If we give a cursory glance at the lower limbs of apes, we see that all
the same characteristics are present in their tarsus that we find in
the human tarsus. In each case there is an astragalus, an _os calcis_,
a scaphoid bone, three cuneiform bones, and a cuboid bone. There are
undoubtedly several peculiarities in which the tarsus differs from
the corresponding part of the human foot. The first metatarsal bone
is joined to the first cuneiform bone by an articular facet which
extends from the back to the sole of the foot. This joint plays a part
resembling that of the thumb of the human hand (see Figs. 20 and 46).
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