Bischoff at first denied the existence of the plantaris in the
chimpanzee, and Brühl had previously done the same, but it is as
normally present in that animal as in man, in whom also it is sometimes
absent. I, however, as well as other observers, have failed to discover
it in the gorilla, orang, and gibbon. The popliteus is developed in
every instance. The tibio-fibular muscle (_Musculus peroneotibialis_),
covered by the popliteus, of which the existence was ascertained by
Gruber, has not been observed by me in any of the anthropoids, with the
exception of the chimpanzee. But it was very apparent in a red sea-cat
monkey (_Cercopithecus ruber_).
The gastrocnemius, which is easily separable into two heads, and the
peroneal muscles have not the same relative width in anthropoids and
man, since in the former case the calf of the lower limb is small, and
it lacks the pleasing roundness which characterizes this part of the
human structure. These muscles, especially in the orang and gibbon,
appear to take a lateral direction. The Tendo Achillis is present,
but it has not the prominent development in height and width which
we observe in man. The long extensor, flexor, and tibial muscles are
in all cases fully developed. The peroneus tertius, as it is termed,
although it should only be regarded as a part of the extensor longus
digitorum, is absent in anthropoids.[74] I myself am not disposed, with
Huxley, Bischoff, and others, to regard this muscle as an abductor.
Brühl perceived in a chimpanzee a fourth rudimentary peroneal muscle
(_Musculus peroneus intermedius_), extending between the peroneus and
the little toe, a muscle sometimes found in man, and which I have
myself only observed in one adult chimpanzee. In the gorilla and the
chimpanzee the extensor longus digitorum passes through a remarkably
strong transverse ligament, formed of fibrous cartilage, which covers
the tarsus. It acts upon the four outer toes (Fig. 55). Brühl has
described the characteristic contraction and extension of the tendons
of the long and short extensors of the toes in the chimpanzee, but I
have myself found some difficulty in producing this action. In Fig. 55
I have endeavoured to represent this condition in the most natural way.
The extensor proprius pollicis is in all cases developed. The extensor
brevis digitorum produces a large, oblique belly for the great toe
(Fig. 55). In the gorilla there is for the great toe an abductor, a
bicipital flexor, an adductor, and an opponens (comp. Fig. 54).
From the extensor brevis digitorum the belly for the great toe rises
with a certain independence. On the right foot of a chimpanzee I
observed a fifth belly of this muscle, going to the little toe (Fig.
55). As my illustration is taken from this specimen, I have represented
the foot with, or in spite of, this interesting anomaly, which, as we
know, sometimes occurs in man.
[Illustration:
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