Appletons' Popular Science Monthly, November 1898: Volume 54, November 1898Various
Science
Appletons' Popular Science Monthly, November 1898: Volume 54, November 1898
Various
Science -- Periodicals; Technology -- Periodicals
=Natural Selection and Fortuitous Variation.=--The three principal
objections urged against Darwin's theory of natural selection were
stated by Prof. W. F. R. Weldon, in his presidential address to the
Section of Zoology and Physiology of the British Association, as being
that the species of animals we know fall into orderly series for the
selection of which purely fortuitous variations can not be supposed
to afford opportunity; that minute structural variations can not be
supposed to affect the death-rate so much as the theory requires they
should, while many of the characters by which species are distinguished
appear to us so small and useless that they can not be supposed to
affect the chance of survival at all; and that the process of evolution
by natural selection is so extremely slow that the time required for its
operation is longer than the extreme limit of time given by estimates of
the age of the earth. The first of these objections the speaker alleged
to be due to a misunderstanding of words; we regard as fortuitous what
we do not understand; and he proceeded to explain how what we call
chance may be shown, especially by a method developed by Professor
Pearson, to be a real and important factor. To the other two objections
Professor Weldon opposed the results of observations of his own and of
Mr. Herbert Thompson on the small shore crabs (_Carcinus mœnas_) at
Plymouth Beach. "In these crabs small changes in the size of the frontal
breadth do, under certain circumstances, affect the death-rate; and the
mean frontal breadth among this race of crabs is, in fact, changing
at a rate sufficiently rapid for all the requirements of a theory of
evolution." In conclusion, he said: "I hope I have convinced you that
the law of chance enables one to express easily and simply the frequency
of variations among animals, and I hope I have convinced you that the
action of natural selection upon such fortuitous variations can be
experimentally measured, at least in the only case in which any one has
attempted to measure it. I hope I have convinced you that the process
of evolution is sometimes so rapid that it can be observed in the space
of a very few years." The whole difficulty of natural selection, he
added, is a quantitative difficulty; and he insisted upon the need of
observations and measurements of the rates of variation.
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