Rupen's son Constantine succeeded him. It was by his help that the leaders of the first crusade captured Antioch. Constantine was succeeded by his two sons, Leo and Theodore jointly, but finally Leo reigned alone; he was an able prince, fought the Saracens with success, and much enlarged his kingdom, and at last made a naval attack on Isaurian Seleucia, the frontier fortress of the Byzantine Empire in this part, and an important seaport. This brought "Handsome John," the ablest of the Comnenian line of Byzantine Emperors, into the field; he stormed the Cilician seaports, and then reduced the chief interior fortresses; Leo fled to the Taurus Mountains, but was captured, and died in captivity at Constantinople. His son Rupen had his eyes put out on a charge of treason, and died of it; but his other son, Toros, escaped, and after John's death restored the Cilician kingdom, which had temporarily been made vassal by John. Toros is the glory of the whole Rupenian line; he was of the first rank, both as a general and a statesman. He scarcely ever suffered a military reverse. He beat the Byzantine armies in campaign after campaign, and the Seljuks as well; under him the new Armenia was almost a match for all its enemies combined, and no one of them dreamed of attacking it single-handed. Levon was another able ruler, who maintained the power and prosperity of the kingdom; he was an ally of the great Emperor Frederick Barbarossa in the Third Crusade, assisted him in capturing Iconium (1190), and both Frederick and the Greek Emperor Alexius III sent him crowns,--the second no great honor, as Alexius was one of the most contemptible of human beings. In Levon's time the capital of the kingdom was Cis, where there is now a great Armenian monastery with rare manuscripts, the residence of a Catholicos. The changes in the extent of the kingdom are very curious; perhaps most curious of all (since the Armenians were always a race of inland and highland farmers, not seamen), the new kingdom was gradually crowded down on the north and lost two-thirds of its territory in that direction, but steadily extended along the coast until it came to include not only all Cilicia but all of old Isauria clear to its western mountain barrier; hundreds of miles of seaboard, from close to Antioch on the one side, to far west of Cyprus on the other, being indeed a strong maritime power. At the end it had lost these western coast extensions, but still had an area larger than that of the Crimea now, a very considerable power to hold the northeast corner of the Mediterranean.