Aspects of plant life; with special reference to the British floraPraeger, R. Lloyd (Robert Lloyd)
Science
Aspects of plant life; with special reference to the British flora
Praeger, R. Lloyd (Robert Lloyd)
Plants -- Great Britain
To explain the massiveness of a tree trunk we have to remember that,
while the cross-section of any structure varies as the square of its
linear dimension, the volume varies as the cube of the same. If we
double the dimensions of a tree, we increase its weight eight times, but
the strength of the trunk is increased only four times. If a tree 100
feet high is supported on a stem 6 feet in diameter, a tree 200 feet
high of the same proportions would need a stem not 12 feet, but over 17
feet in diameter, to be supported equally efficiently. This proportion
increases rapidly: a similar tree 300 feet high would need a stem 30
feet in diameter; a tree 1,000 feet high would require a stem 180 feet
in diameter, or 32,400 square feet in cross-section. We see, then, why a
limit of tree growth is rapidly reached, at about 300 feet, and why the
trees which grow to that height have trunks which are one of the wonders
of the world, exceeding 30 feet in diameter, or about 100 feet in
circumference.
Climbing stems represent efforts on the part of plants to economize
material by utilizing the rigidity of neighbouring plants, and by
reaching to the light on their shoulders. Here, as in aquatics, the
_rope_ type of stem is in evidence; it resembles a garden hose, offering
great flexibility and conducting capacity, but without rigidity to
support its own weight, much less that of the leaves and flowers which
it bears. To secure support, the stem itself (or branches of it), the
leaves, or the stipules (leafy projections on either side of the
junction of leaf and stem), are used. Sometimes support is obtained by
twining (compare Convolvulus, Grape-Vine, Vetch), sometimes by adherent
discs (Virginia Creeper), or aerial roots (Ivy), often by mere
scrambling, often aided by reflexed hooks on leaf and stem (Bramble,
Cleavers). The mechanism by which twining is accomplished is of great
interest. It is an effect of unequal growth of the different sides of
the stem. If the unequal growth were confined to one side, the stem
would eventually form a coil, or series of circles. But the region of
greatest growth keeps shifting round the stem, with the result that the
tip of the shoot describes a circle or ellipse, like the hand of a clock
pointing successively in all directions. The stimulus is due, as in the
case of the erect growth of ordinary stems (which usually display
similar movements in a less degree) to gravity. Sometimes the movement,
or _nutation_, is in the same direction as that of the hands of a clock
(_e.g._, in the Hop); more frequently it is in the opposite direction,
as of a clock-hand moving backwards. The result of this movement is that
if the shoot encounters, say, an upright stem, it will lap round it in a
spiral manner, and unless the said stem be quite smooth and unbranched,
the twining shoot will be eventually supported by it. How effective the
twining habit is as regards economy of building material may be seen
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