Aspects of plant life; with special reference to the British floraPraeger, R. Lloyd (Robert Lloyd)
Science
Aspects of plant life; with special reference to the British flora
Praeger, R. Lloyd (Robert Lloyd)
Plants -- Great Britain
colouring matter dissolved in the cell-sap (pinks and blues) or to small
coloured solid bodies (_chromoplasts_) contained in the cells (reds and
yellows). Chlorophyll being absent, the coloured petals do not assist
assimilation: they are purely advertisements, though incidentally they
often fulfil a useful protective rôle for the important organs which
they surround. In this latter connection their sensitiveness to changes
of light and temperature, which causes them to close in dark or cold
weather, is a very familiar phenomenon; as is also the excellent
protection which they provide in flowers such as those of the _Labiatæ_,
where, fused together into a tube, they form a kind of cave in which the
stamens and pistil nestle securely.
[Illustration: FIG. 24.--FRUIT OF CORIARIA JAPONICA. 1/1.]
An exceptional use of petals, where indeed they are used for the
purposes of advertisement, but to secure the dispersal not of the
pollen, but of the seeds, is illustrated in Fig. 24. In the genus
_Coriaria_ the staminate and pistillate organs are borne on separate
flowers. The flowers of both kinds are small and inconspicuous. But in
the “female” flowers the petals persist after flowering, and, becoming
fleshy and comparatively large, enclose the seed in a pulpy berry-like
envelope, which no doubt serves the same purpose as a true berry in
securing seed-dispersal by being devoured by birds. In _C. terminalis_,
which comes from the Himalayas, the “ripe” corolla is bright orange; in
_C. japonica_, from Japan, it is at first coral-red, and when mature
velvet-black.
The _stamens_, which form the next ring (sometimes a double ring or a
close spiral), are much less leaf-like than the sepals or petals, yet
there can be no doubt that they are descended from leaf-shaped organs;
this is especially clear from the study of certain primitive fossil
types, in which the corresponding organs which bear the pollen are
actually leaf-like. In most of the present-day Seed Plants the stamens
conform to a uniform type--a slender stalk (_filament_) bearing a head
(_anther_) containing four chambers, in which are produced _pollen
grains_, which escape when the flower is mature by the splitting of the
enclosing walls. The ways in which the pollen is then conveyed to the
pistil of other flowers have been referred to briefly on a previous page
(p. 82). The stamens in many flowers are few, and their number usually
bears a relation to the number of the other floral parts; in other
flowers, for instance Rose and St. John’s wort (_Hypericum_), they are
of large and indefinite number. The peculiar arrangement of the pollen
in Orchids has been already noted (p. 94).
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