Contemporaneous with Zeuxis, and equal in fame, was Parrhasius, a native of Ephesus, whose skill lay in accuracy of drawing and power of expression. He gave to painting true proportion, and attended to minute details of the countenance and the hair. In his gods and heroes, he did for painting what Phidias did in sculpture. His outlines were so perfect as to indicate those parts of the figure which they did not express. He established a rule of proportion which was followed by all succeeding artists. While many of his pieces were of a lofty character, some were demoralizing. Zeuxis yielded the palm to him, since Parrhasius painted a curtain which deceived his rival, whereas the grapes of Zeuxis had deceived only birds. Parrhasius was exceedingly arrogant and luxurious, and boasted of having reached the utmost limits of his art. He combined the magic tone of Apollodorus with the exquisite design of Zeuxis and the classic expression of Polygnotus. Many were the eminent painters that adorned the fifth century before Christ, not only in Athens, but in the Ionian cities of Asia. Timanthes of Sicyon was distinguished for invention, and Eupompus of the same city founded a school. His advice to Lysippus is memorable: "Let Nature, not an artist, be your model." Protogenes was celebrated for his high finish. His Talissus took him seven years to complete. Pamphilus was celebrated for composition, Antiphilus for facility, Theon of Samos for prolific fancy, Apelles for grace, Pausias for his chiaro-oscuro, Nicomachus for his bold and rapid pencil, Aristides for depth of expression.