The formation of resting-spores of vaucheria is accomplished by means
of special organs, _oögonia_ (_o_, Fig. 268) and _antheridia_ (_a_,
Fig. 268). These are both specially developed branches from the
thallus. The antheridia are nearly cylindrical, and curved toward
the oögonia. The upper part of an antheridium is cut off by a cross
wall, and within it numerous ciliated _sperm-cells_ are formed. These
escape by the ruptured apex of the antheridium. The oögonia are more
enlarged than the antheridia, and have a beak-like projection turned
a little to one side of the apex. They are separated from the thallus
thread by a cross wall, and contain a single large green cell, the
_egg-cell_. The apex of the oögonium is dissolved, and through the
opening the sperm-cells enter. Fertilization is thus accomplished.
After fertilization the egg-cell becomes invested with a thick wall
and is thus converted into a resting-spore, the _oöspore_.
[Illustration: FIG. 269.—FUCUS. Fruiting branches at _s_, _s_. On the
stem are two air-bladders. ]
* * * * *
_Fucus._—These are rather large specialized algæ belonging to the
group known as brown seaweeds and found attached by a disk to the
rocks of the seashore just below high tide (Fig. 269). They are firm
and strong to resist wave action and are so attached as to avoid
being washed ashore. They are very abundant algæ. In shape the plants
are long, branched, and multicellular, with either flat or terete
branches. They are olive-brown. Propagation is by the breaking off of
the branches. No zoöspores are produced, as in many other seaweeds;
and reproduction is wholly sexual. The _antheridia_, bearing
_sperm-cells_, and the _oögonia_, each bearing eight _egg-cells_,
are sunken in pits or _conceptacles_. These pits are aggregated in
the swollen lighter coloured tips of some of the branches (_s_, _s_,
Fig. 269). The egg-cells and sperm-cells escape from the pits and
fertilization takes place in the water. The matured eggs, or spores,
reproduce the fucus plant directly.
* * * * *
[Illustration: FIG. 270.—NITELLA.]
_Nitella._—This is a large branched and specialized fresh-water alga
found in tufts attached to the bottom in shallow ponds (Fig. 270).
Between the whorls of branches are long _internodes consisting of a
single cylindrical cell_, which is one of the largest cells known in
vegetable tissue. Under the microscope the walls of this cell are
found to be lined with a layer of small stationary chloroplastids,
within which layer the protoplasm, in favourable circumstances,
will be found in motion, moving up one side and down the other (in
rotation). Note the clear streak up the side of the cell and its
relation to the moving current.
FUNGI
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