When the mycelium is about two days old, it begins to form the
long fruiting stalks which we first noticed. To study them, use a
compound microscope magnifying about two hundred diameters. One
of the stalks, magnified, is shown in _a_, Fig. 274. It consists
of a rounded head, the _sporangium_, _sp_, supported on a long,
delicate stalk, the _sporangiophore_. The stalk is separated from the
sporangium by a wall which is formed at the base of the sporangium.
This wall, however, does not extend straight across the thread, but
it arches up into the sporangium like an inverted pear. It is known
as the _columella_, _c_. When the sporangium is placed in water,
the wall immediately dissolves and allows hundreds of spores, which
were formed in the cavity within the sporangium, to escape, _b_.
All that is left of the fruit is the stalk, with the pear-shaped
columella at its summit, _c_. The spores that have been set free by
the breaking of the sporangium wall are now scattered by the wind and
other agents. Those that lodge in favourable places begin to grow
immediately and reproduce the fungus. The others soon perish.
[Illustration: FIG. 274.—MUCOR.
_a_, sporangium; _b_, sporangium bursting; _c_, columella.]
The mucor may continue to reproduce itself in this way indefinitely,
but these spores are very delicate and usually die if they do not
fall on favourable ground, so that the fungus is provided with
another means of carrying itself over unfavourable seasons, as
winter. This is accomplished by means of curious _thick-walled
resting-spores_ or _zygospores_. The zygospores are formed on the
mycelium buried within the substance on which the plant grows. They
originate in the following way: Two threads that lie near together
send out short branches, which grow toward each other and finally
meet (Fig. 273). The walls at the ends, _a_, then disappear, allowing
the contents to flow together. At the same time, however, two other
walls are formed at points farther back, _b_, _b_, separating the
short section, _c_, from the remainder of the thread. This section
now increases in size and becomes covered with a thick, dark brown
wall ornamented with thickened tubercles. The zygospore is now mature
and, after a period of rest, it germinates, either producing a
sporangium directly or growing out as mycelium.
[Illustration: FIG. 273.—MUCOR, showing formation of zygospore on the
right; germinating zygospore on the left. ]
The zygospores of the mucors form one of the most interesting and
instructive objects among the lower plants. They are, however, very
difficult to obtain. One of the mucors (_Sporodinia grandis_) may be
frequently found in summer growing on toadstools. This plant usually
produces zygospores that are formed on the aërial mycelium. The
zygospores are large enough to be recognized with a hand lens. The
material may be dried and kept for winter study, or the zygospores
may be prepared for permanent microscopic mounts in the ordinary way.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account