Numerous _summer-spores_ are formed of short, erect branches all over
the white surface. One of these branches is shown in Fig. 277. When
it has grown to a certain length, the upper part begins to segment or
divide into spores which fall and are scattered by the wind. Those
falling on other willows reproduce the fungus there. This process
continues all summer, but in the later part of the season provision
is made to maintain the mildew through the winter. If some of the
white patches are closely examined in July or August, a number of
little black bodies will be seen among the threads. These little
bodies are called _perithecia_, shown in Fig. 278. To the naked eye
they appear as minute specks, but when seen under a magnification of
200 diameters they present a very interesting appearance. They are
hollow spherical bodies decorated around the outside with a fringe
of crook-like hairs. The _resting-spores_ of the willow mildew are
produced in sacs or _asci_ inclosed within the leathery perithecia.
Figure 279 shows a cross-section of a perithecium with the asci
arising from the bottom. The spores remain securely packed in the
perithecia. They do not ripen in the autumn, but fall to the ground
with the leaf, and there remain securely protected among the dead
foliage. The following spring they mature and are liberated by the
decay of the perithecia. They are then ready to attack the unfolding
leaves of the willow and repeat the work of the summer before.
* * * * *
_The wheat rust._—The development of some of the rusts, as the
common _wheat rust_ (_Puccinia graminis_), is even more interesting
and complicated than that of the mildews. Wheat rust is also a true
parasite, affecting wheat and a few other grasses. The mycelium
here cannot be seen by the unaided eye, for it consists of threads
which are present within the host plant, mostly in the intercellular
spaces. These threads also send short branches, or _haustoria_ (Fig.
132), into the neighbouring cells to absorb nutriment.
[Illustration: FIG. 280.—SORI CONTAINING TELEUTOSPORES OF WHEAT RUST. ]
[Illustration: FIG. 281.—TELEUTOSPORE OF WHEAT RUST. ]
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