Biology and Its Makers: With Portraits and Other IllustrationsLocy, William A. (William Albert)
History
Biology and Its Makers: With Portraits and Other Illustrations
Locy, William A. (William Albert)
Biology -- History
Passing over his book, _The Germ Plasm_, published in English in 1893,
we may fairly take his last book, _The Evolution Theory_, 1904, as the
best exposition of his conclusions. The theoretical views of Weismann
have been the field of so much strenuous controversy that it will
be well perhaps to take note of the spirit in which they have been
presented. In the preface of his book just mentioned, he says: "I make
this attempt to sum up and present as a harmonious whole the theories
which for forty years I have been gradually building up on the basis
of the legacy of the great workers of the past, and on the results of
my own investigations and those of my fellow-workers, not because
I regard the picture as incomplete or incapable of improvement, but
because I believe its essential features to be correct, and because
an eye-trouble which has hindered my work for many years makes it
uncertain whether I shall have much more time and strength granted to
me for its further elaboration."
The germ-plasm theory is primarily a theory of heredity, and only when
connected with other considerations does it become the full-fledged
theory of evolution known as Weismannism. The theory as a whole
involves so many intricate details that it is difficult to make a clear
statement of it for general readers. If in considering the theories of
Lamarck and Darwin it was found advantageous to confine attention to
salient points and to omit details, it is all the more essential to do
so in the discussion of Weismann's theory.
In his prefatory note to the English edition of _The Evolution Theory_
Thomson, the translator, summarizes Weismann's especial contributions
as: "(1) the illumination of the evolution process with a wealth of
fresh illustrations; (2) the vindication of the 'germ-plasm' concept
as a valuable working hypothesis; (3) the final abandonment of any
assumption of transmissible acquired characters; (4) a further analysis
of the nature and origin of variations; and (5), above all, an
extension of the selection principle of Darwin and Wallace, which finds
its logical outcome in the suggestive theory of germinal selection."
Continuity of the Germ-Plasm.--Weismann's theory is designated that
of continuity of the germ-plasm, and in considering it we must first
give attention to his conception of the germ-plasm. As is well known,
animals and plants spring from germinal elements of microscopic size;
these are, in plants, the spores, the seeds, and their fertilizing
agents; and, in animals, the eggs and the sperms. Now, since all
animals, even the highest developed, begin in a fertilized egg, that
structure, minute as it is, must contain all hereditary qualities,
since it is the only material substance that passes from one generation
to another. This hereditary substance is the germ-plasm. It is the
living, vital substance of organisms that takes part in the development
of new generations.
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