From these observations it would appear that the roots of most of
the plants growing on the plain have a well-developed main root,
and that few, if any, perennial roots lie near the surface of
the ground. This condition is rather different from that seen
at Laghouat, where the root-system of typical inhabitants of a
similar habitat is of the strictly generalized type and may point
to a difference in some character of the habitat—for instance,
the precipitation at Ghardaia.[24]
In certain portions of the floor of the M’Zab Valley the soil
conditions favor full and normal development of all forms of
root-systems, but in other portions, because of the presence of an
impervious hardpan, such development is not possible. Where there
is hardpan the presence of species with obligate tap-roots is
precluded, while such as have a more flexible root-system (e.g.,
_Peganum harmala_) can to a degree accommodate themselves to
the unfavorable soil conditions. However, one instance was seen,
which will be reported below, where a plant with a tap-root was
found growing on a hardpan stratum so hard as to be excavated with
the greatest difficulty. The character of the root is very greatly
modified by the presence of the hardpan. Following is a sketch of
the root-systems of a few plants growing naturally in the valley.
_Peganum harmala_ is one of the most characteristic plants of
the floor of the M’Zab Valley; so far as my observations go, it
usually occurs where there is much soil and where the water relations
are the most favorable. Several studies of it were carried out on
plants growing in different parts of the valley. In order to make
the leading study of its roots, a typical habitat was selected east
of Ghardaia and nearly in the midst of the valley. Here the upper
soil, to a depth of about 20 cm., is a fine sand carrying water-worn
pebbles and coarse stones. Below the sand is a denser stratum a few
centimeters in thickness, but not the hardpan described in another
place. Below the harder stratum, to an undetermined depth, are
mingled sand and gravel coarser in texture than the superficial sand.
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