Up to this point the plants whose roots have been described were
found growing where the soil was favorable to a fairly normal
development, but the soil of the valley is not all of this character,
and where hardpan is present the substratum is often extremely
hard and, one would think from inspection, impermeable to water
as well as impenetrable to roots. In such habitats the number of
individuals and also of species is very naturally limited.
About 2 kilometers west of Ghardaia is an area where the hardpan
reaches nearly or quite to the surface. Here _Peganum harmala_
is to be found, since its generalized type of root-system is
capable of not a little variation, adjusting the species to a
variety of soil conditions not otherwise possible. Associated with
_Peganum_ was a single specimen, of dwarfed and badly eaten form,
growing out of the hardpan itself. The position of the plant was
so unusual that its root-system was in part excavated. The form,
an undetermined chenepod, was found to have the exceptional form of
root, thus proving the rule just suggested, for it had a well-marked
tap-root. A gnarled main root was found to take a zigzag course
through 27 cm. of hardpan to the softer stratum beneath. The root
did not follow a crack, but struck boldly downward through soil so
hard that it was removed by the use of a sharp iron instrument and
only with great difficulty.
LEAF-HABITS IN THE GHARDAIA REGION.
Observations of the leaf characters of the desert perennials, as
shown in November, offer some points of interest, especially since
there had been no rain for a year previous to the visit. As would be
expected, the leaf-habit is various, ranging from scale-like or none
to fair size. Several species appear to be evergreen, including
_Artemisia herba-alba_, _Capparis spinosa_, _Echium humilis_,
_Fagonia bruguieri_, _Gymnocarpon_ sp., _Haloxylon articulatum_,
_Helianthemum sessiliflorum_, _Henophyton deserti_, _Herniaria
fruticosa_, and _Salsola_. Certain plants probably, as _Henophyton_,
are facultative evergreens, retaining the leaves if the season is
moist and dropping them if it is excessively dry; several of this
genus were seen without leaves, but with green stems. (See, also,
figs. 39, 40, and 41.)
Some idea of the variation in size of the leaves of a single species,
as well as the average size of the leaves, will be had from the
results of a few measurements. The leaves of _Henophyton deserti_
range in length from 2 to 3.1 cm., and in breadth from 0.2 to 0.5
cm. The average length of 12 leaves from a single branch 7 inches
in length was found to be 2.82 cm.; the average breadth was 0.35
cm. On another branch, 23 cm. long, were 35 mature leaves which
averaged almost exactly the size of those just given. Since these
leaves are rather numerous and of good size, it will appear at once
that the species has a relatively large leaf-surface, even if the
area can not be stated more definitely. (Fig. 30.)
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