reason why such temporary rootlets should not be formed nearer the
surface. However, should there be sufficient moisture in the soil
to permit absorption by roots, provided a slow rate was adequate
to replace the transpiration loss, which rate was made lower by a
more moist air, a slight rain would be of great significance, even
if it did not penetrate to any appreciable depth. On the intensely
arid desert such slight modifications of the water relations as the
lowering of the temperature as winter approaches, causing decreased
evaporation or rains, although actually small in amount, may be of
large moment to plants. Such a condition was noted at Ghardaia,[35]
where there had been a drought for over twelve months, but on the
return of the cool season, with a lower evaporation rate, growth
was resumed and several plants came into flower. This appeared to be
not wholly the stimulus of lower temperature, since I was informed
by good authority that the plants renewed their various activities
whenever rains chanced to come, whatever might be the season.
From the observations last given it appears that sufficient moisture
persists in the soil to tide perennials over the long periods of
drought, although not in sufficient amount to permit active growth
during the dry seasons. This is not an uncommon occurrence with
desert plants. For example, whenever in the Tucson region the arid
seasons are uncommonly long, or there has been a relatively small
rain, as 25 per cent less than usual, much of the vegetation may
remain dormant. Under such conditions an evergreen like _Covillea
tridentata_ drops all save the youngest and smallest leaves and
maintains this nearly defoliate condition for long periods, with
little other change. The plant behavior noted is always connected
with insufficient moisture.
The soil in the oueds and probably also in the reg, as well as
that of the dayas, is of considerable depth. The soil depth, or
rather the depth before solid rock is encountered in the valley
of the Oued M’Zab at Ghardaia, is from 20 to 30 meters, or even
more; at the Daya Tilrempt the depth to water in one of the wells
is about 95 meters. Although there is much gravel and sand in the
fill of these depressions, it is likely that they afford the most
favorable soil conditions in the desert for the development of
a large root-system. Be that as it may, it is certain that large
plants, such as _Tamarix_ and _Pistacia_, occur only in such places.
Where there is most soil on the hamada, as in certain pockets near
the old hill town of Ghardaia, abandoned several centuries since,
one finds also the most and largest plants of this, the hamada,
formation. Moreover, at the time in November when growth was noticed
in the oued plants, it was also taking place in these favoring
situations, about 50 meters above the level of the valley of the
M’Zab and much above any possible permanent water-supply such as
is afforded by a water-table.
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