Nor was the earth peopled wholly by this ancestor of _Lycopodium
Selago_, for we find at this time, or just after it, a great development
of plants of this type. Some of these were giant, treelike club-mosses
that have been so well preserved as fossils that even their internal
structure and spore-bearing characters are well known. Many other
strange relatives of our modern club mosses flourished in those days,
some of which have wholly disappeared, as have all the treelike forms.
These highly organized club mosses, quite unlike any modern
representative of the family, appear to have been crowded off the earth
by other and subsequent types of vegetation, while _Lycopodium Selago_,
and about thirty related species, have persisted to the present day; not
precisely in all cases as they were in this dawn of a land flora, but in
many cases with modern structure and reproductive processes so close to
the ancestral types as to be nearly identical.
Perhaps nothing gives one a better impression of the tremendous time
that must have elapsed before the appearance of these ancient club
mosses than the very slight modifications from their ancient condition
which their structure at the present time exhibits. While nothing is
certainly known of their origin, when they first appeared they were
plants with a well-developed vascular system, having stems and leaves
quite unlike any of their predecessors’, and a reproductive process
almost precisely like their modern descendants. In other words, if they
have changed so slightly in all the millions of years since our
rock-written records of them first occur, what an infinitely greater
period must have elapsed down the dim vista of the ages before their
appearance. Of this period, with the exception of fossil algæ, we know
practically nothing, and, worst of all, the actual transition from a
wholly water-inhabiting flora to these certainly land-inhabiting club
mosses may never be known. For, added to the difficulty of water plants
being preserved as fossils, already mentioned, is the fact that as they
are the oldest, they are found in the deepest strata and, consequently,
the hardest to find; and due to changes in the earth’s crust, these
ancient fossil-bearing strata have often been much disturbed.
The conclusion appears to be indicated that the origin of a land flora
came about with the appearance of these ancient club mosses, which are
not mosses in our present-day interpretation of those plants, and that
at about the same period many other plants also were found, the whole
vegetation resembling nothing that exists at the present time, but many
of the different kinds of this ancient flora showed unmistakable
evidence of being the progenitors of many plants that exist to-day. What
these were, and particularly what they accomplished, both in the history
of the plant kingdom and in making the world habitable for man who did
not come for millions of years after they were preparing the way, will
be considered in
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