One of the main facts about variation is that it is itself a very
variable thing, and the nature of those “fluctuating variations” which
are so common in nature well illustrate the point. Within what we know
as a species there are many individuals that vary one way or the other
from a fairly central, we might almost call them normal, mass of
individuals which are typical of that species. Nearly all these forms on
the fringe of the species due to the environmental changes and not to
changes of hereditary constitution will, if left to themselves, tend
through their progeny to become more like the central mass as time goes
on, while their position, or some other equally nontypical edge of the
fringe, will be taken up by other variants from the average conditions.
The amount of this fluctuating variability among plants is beyond
calculation, and its action has often been likened to the swinging of a
pendulum, which of course spends twice the time passing through the
center of its arc, that it does on the limits of it. This very nearly
expresses the proportion of fluctuating variants to the mass of typical
individuals in many species of plants. In many others, often peculiarly
unstable species, the number of individuals at the fringe is very large
indeed. Sometimes there may be one or more that do hold their
characteristics, in which case we know that they are not true
environmental variations but have actually a different constitution.
These will be considered presently under another and different sort of
variability. But, speaking generally, these fluctuating or environmental
variants are merely forms of the species, and, other things being equal,
they will not actually originate new species.
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