“Now we may see what this amounts to. We have got the same genus of
fishes represented in a variety of specific types from the Sea of
Galilee and the Jordan that are found in the feeders of the Nile,
and in the Central African lakes down to the Zambezi. The conclusion
is natural that all these fishes come from a common origin, and that
during the Tertiary period there was a chain of fresh-water lakes,
extending to the lakes in Africa, similar to the chain of lakes in
North America.
“We find in Palestine forty-three species of fishes, of which only
eight belong to the ordinary ichthyological fauna of the Mediterranean
rivers. But these belong to the rivers of the coast. In the Jordan
system only one species out of thirty-six belongs to the ordinary
Mediterranean fauna, viz., _Blennius lupulus_. Two others, _Chromis
niloticus_ and _Clarias macracanthus_, are Nilotic. Seven other
species occur in other rivers of South-Western Asia, the Tigris,
Euphrates, &c. Ten more are found in other parts of Syria, chiefly in
the Damascus lakes, and the remaining sixteen species of the families
_Chromidæ_, _Cyprinodontidæ_, and _Cyprinidæ_, are peculiar to the
Jordan, its affluents, and its lakes. This analysis points at once to
the close affinity of the Jordan with the rivers of Tropical Africa.
The affinity is not only of species, but of genera, for _Chromis_
and _Hemichromis_ are peculiarly Ethiopian forms, while the other
species are identical with, or very closely allied to, the fishes
from other fresh waters of Syria. But the African forms are a very
large proportion of the whole, and considering the difficulty of
transportation in the case of fresh water fishes, the peculiarities of
this portion of the fauna are of great significance.
“The fluviatile fishes claim special attention, dating, as they
probably do, from the earliest time after the elevation of the country
from the Eocene ocean. In the _Foraminifera_, mentioned above as found
in the Dead Sea sand, such as _Gr. capreolus_, we have the relics of
the inhabitants of that early sea. But of the living inhabitants we
must place the Jordanic fishes as the very earliest, and these, we have
seen, form a group far more distinct and divergent from that of the
surrounding region than in any other class of existing life. During
the epochs subsequent to the Eocene, owing to the unbroken isolation
of the basin, there have been no opportunities for the introduction
of new forms, nor for the further dispersion of the old ones. These
forms, as we have seen, bear a striking affinity to those of the
fresh-water lakes and rivers of Eastern Africa, even as far south as
the Zambezi. But the affinity is in the identity of genera, _Chromis_
and _Hemichromis_ being exclusively African, while the species are
rather representative than identical.
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