The latter is perhaps the case with the curious little Rodent which
alone forms the present family, of which its original describer,
M. Alphonse Milne-Edwards, writes as follows:--“In its general
aspect it somewhat resembles certain Opossums, and like these it is
pedimanous;[52] but these are the only analogies it presents to the
Marsupials, and in its dental system, as also in the rest of its
organisation, we easily see that it belongs to the order Rodentia. It
differs, however, from all the members of this group by characters of
considerable importance; I may even say that, by some peculiarities
of structure it departs from all other Mammals, and that we find in
it anatomical arrangements of which we have hitherto had examples
only in the class of Reptiles.” After an exhaustive discussion of the
characters of this curious little animal, M. Milne-Edwards comes to
the conclusion that it is most nearly related to the members of the
following family, and especially to the Hamsters, although he found it
impossible to unite it with them. In this course he has been followed
by other writers.
[Illustration: LOPHIOMYS.]
The general construction of the skull is the same as in the Muridæ,
but from the temporal ridges thin plates are developed, which bend
downwards, and articulate with similar plates springing from the malar
bones, and thus completely arch over the temporal fossæ after a fashion
only met with in certain reptiles, and especially in the Hawksbill
Turtle (_Chelone caretta_). The whole upper surface of the skull is
covered with minute but perfectly definite granules, arranged with much
regularity, and these, which occur in no other Mammal, give the skull
a very peculiar aspect, such as may be seen in some fishes. As in the
Muridæ, there are three molars on each side in each jaw, and these are
rooted and strongly tubercular; the foremost in each series having
three and the others each two ridges. Without entering in detail into
the peculiarities described at great length by M. Milne-Edwards, we
may say that in its general structure, and especially in that of the
skeleton, the animal is murine, but with a very important distinction,
namely, that the collar bones, which are well developed in the Rats and
their allies, are here reduced, as in the Hares and Rabbits, so as to
form only two small bony styles freely suspended among the muscles,
and that the first toe in the hind feet, although not very long, is so
attached as to be opposite to the rest, thus converting the organ into
a prehensile hand which the animal uses freely in climbing. The cæcum
is small.
Public-domain text, read in full here on John Shaqi.
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