This second family of the section Hystricomorpha exhibits the
conversion of the hairs into spines in perfection, the whole upper part
of the body being in several instances completely covered with long,
hollow, pointed quills, whilst in all cases great numbers of spines
and stiff bristles are mixed with the hair. The form of the skull in
these animals is distinctive. It is ovate, the cranial portion being
more or less inflated by air-cavities in the bones, and the facial
portion short, but the occipital or hinder surface is usually nearly
perpendicular; the malar portion of the zygoma has no angular process
as in the preceding family; the molar teeth are four in number on each
side in each jaw; and the limbs are about equal in development. The
incisor teeth are large and powerful. With regard to the development of
the tail there are considerable differences, some species having that
organ quite short, while in others it is of moderate length, or long
and sometimes prehensile.
[Illustration: SKULL OF THE PORCUPINE.]
The Porcupines fall readily into two distinct groups (sub-families)
characterised by structure, habits, and geographical distribution. In
the strictly terrestrial species, or True Porcupines (HYSTRICINÆ),
which inhabit the warmer parts of the eastern hemisphere, the skull
is rather more elongated than in the others; the front margin of the
orbit is over the third molar; the molars are rootless when young, but
become closed after a time, and the clavicles are imperfect. The upper
lip is furrowed; the tail, which may be either long or short, is never
prehensile; the soles of the feet are smooth; and the female has six
teats.
[Illustration: COMMON PORCUPINE.]
The arboreal species (SPHINGURINÆ), which are all American, have the
skull peculiarly short, the front margin of the orbit over the first
molar, the molars always rooted, and the clavicles perfect. The upper
lip is not furrowed; the tail is moderate or long, and generally
prehensile; the soles of the feet are covered with wart-like tubercles;
and the female has only four teats.
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