The CASTORIDÆ, including at present only a single species common to
the northern parts of both hemispheres, are represented by several
peculiar fossil forms. Remains of the Common Beaver (_Castor fiber_)
are not uncommon in peat bogs and other late superficial deposits both
in Europe and America; and, according to Sir R. Owen, in association
with those of the Rhinoceros, Mammoth, and Mastodon, even in the
Fluvio-marine Crag (Newer Pliocene) of Norfolk. In Belgium its bones
have been found in caves. Among the Mammals from the Upper Tertiaries
of the Sivalik Hills, Messrs. Falconer and Cautley record a Beaver
distinct from the existing species, although nearly allied to it.
The skull of a great Beaver, one-fifth larger than that of the
living species, was obtained many years ago by M. Fischer from sandy
deposits on the shores of the Sea of Azov, and, as it differed in
some peculiarities of the teeth from _Castor fiber_, was described by
him as constituting a distinct genus under the name of _Trogontherium
Cuvieri_. It is now regarded as a true Beaver, and named _Castor
Trogontherium_. The British species, described and figured by Sir R.
Owen from the Norfolk Forest bed under the name of _Trogontherium
Cuvieri_, is, however, quite distinct, and belongs to the genus
_Diobroticus_, characterised by having the third upper and first lower
molar teeth with four enamel folds, and the rest only with two, most of
the folds soon becoming isolated as the teeth wear down. This animal
must have been nearly twice the size of the European Beaver.
At a still earlier period--namely, in the Miocene--the family Castoridæ
was represented, both in Europe and America, by some small species,
nearly agreeing with the Beavers in dentition, but differing in
the characters of certain bones of the skull. These form the genus
_Steneofiber_. The largest (_S. viciacensis_), from the Miocene of
the Allier, was about half the size of the Beaver; another (_S.
sansaniensis_), from the fresh-water limestone of Sansan, was about as
large as a Rat; an American species (_S. nebrascensis_), from the “Bad
Lands” of Dakota, was rather smaller than a Marmot, and presented some
resemblance to the Agoutis in the characters of the teeth; and a fourth
species (_S. pansus_) occurs in the Santa Fé marls. _Eucastor tortus_,
a species rather smaller than a Marmot, is described by Dr. Leidy as
very nearly related to the Beaver. Its remains were found in loose
sands of the Niobrara River, Wyoming. _Chalicomys_ and _Palæomys_ are
genera doubtfully placed here. Their species occur in the Miocene and
Pliocene of Europe.
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