No fossil LOPHIOMYIDÆ have yet been detected, but the great family
MURIDÆ has left abundant evidence of its former existence. Species of
the genera _Mus_, _Arvicola_, _Myodes_, and _Cricetus_, identical in
many cases with those now living, have been obtained frequently in
Post-Pliocene deposits and in bone-caves in Europe. Lemmings (_Myodes
lemmus_ and _torquatus_) are recorded from English caves. The genus
_Mus_ is also represented by several species in the Miocene deposits of
France, and in the Sivalik beds investigated by Falconer and Cautley.
The Miocene of Sansan has furnished a form which has been doubtfully
regarded as a Gerbille, and named _Meriones Laurillardi_. In the same
and other deposits of the same age in South-eastern France several
species of an extinct genus (_Cricetodon_) have been obtained. Their
dentition resembles that of the Hamster, but the first molars in both
jaws have a tubercle less; the largest species (_C. sansaniensis_)
rather exceeded the Hamster in size, while the smallest was less
than a Mouse. Associated with some of these are two doubtful forms,
_Decticus_ and _Elomys_, the latter considered by M. Aymard, its
describer, to be allied to _Hydromys_. The American fossil Muridæ are
for the most part either species of the genus _Hesperomys_, or nearly
related to it. Twelve species of that genus were obtained by Dr. Lund
from the Brazilian bone-caves, but of these eight were identified by
him with species still existing. In North America two species of a
nearly-allied genus (_Eumys_) have been obtained from Miocene deposits;
and the bone-caves of Pennsylvania furnish the remains of a species of
_Neotoma_ (_N. magister_), hardly distinguishable from the Florida Rat.
A _Rhizomys_ from the Sivalik deposits of North-western India is the
only recorded fossil representative of the SPALACIDÆ; and of the
GEOMYIDÆ the only known species are a _Geomys_ from the Pliocene of
Nebraska, nearly allied to, if not identical with, the living _G.
bursarius_; and one from the “Tertiaries of the Plains,” described by
Professor Cope as _Colotaxis cristatus_, which, however, has only three
molars in the lower jaw.
The DIPODIDÆ are still more scantily represented. A Jerboa described by
M. Fischer from Post-Pliocene deposits, probably of Tartary, is very
nearly allied to the living _Dipus platurus_, but has shorter toes
and broader cannon-bones. The genus _Dipoïdes_, from the “Bohnerz” of
Würtemberg, is founded on a single tooth, and its position in this
family is very doubtful.
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