After the age of the Trias, when there was much continuous land
surface, Europe was broken up into a coral island tract, during the age
of the collection of the Jurassic deposits. The islands were tenanted
by many small Marsupials, four species of which have been discovered in
the deposits of Stonesfield slate at the bottom of the Great Oolite.
They belong to the extinct genera Amphitherium, Phascolotherium,
and Stereognathus, and the first somewhat resembled the Myrmecobius
of recent times; but all that can be said is that they belonged to
Marsupial animals. Piled on the Stonesfield slates are many hundred
feet of strata, and high up amongst them, in the Swanage and Purbeck
districts, are deposits in which Messrs. Brodie and Beckles have found
portions of the skeletons of numerous insectivorous Marsupials, of
which the genera Spalacotherium, Plagiaulax, Triconodon, and Galestes
are the most important. They were small, as a rule, and there has been
much debate regarding their affinities with modern insectivorous forms,
and they are still surrounded with doubt.
The appearance of the Mammalia without pouches took place in the
Eocene age, and in the Old and New World, and contemporaneously with
them lived in France a kind of Opossum, some of whose bones were
found in the strata of Montmartre, near Paris; and in later Tertiary
strata other relics have been found. These are the only instances of
a fossil Didelphid occurring out of the New World; and there, where
the Opossums are now characteristic animals, they were present in the
last geological age, for in the Brazilian latest deposits remains of
several species of Didelphys have been found. Remains of these fossil
Opossums have been found in the North American Pliocene deposits.
The more ancient deposits of Australia have not yielded the remains
of any of the animals which are now so peculiar to the province, but
in the bone caves of the Wellington Valley, some two hundred and ten
miles west of Sydney, Sir Thomas Mitchell discovered a mass of bones,
forming a breccia with limestone, which contained numerous and most
interesting Marsupial remains. In deposits of the same late age, and
in bogs and gravels in Queensland, other remains were found. They were
described by Sir R. Owen in one of his greatest works, and they belong
to the Australian families of Marsupials, and not to the American
Didelphidæ. As was usual elsewhere before the appearance of man on the
earth, and contemporaneously with him for awhile, many of the kinds
which resemble more or less those now living, or would be classified
in the same family, and perhaps in the same genus, are gigantic. Owen
distinguished among the bones those of large fossil Marsupials which
belong to the Macropodidæ, and which may be arranged as subdivisions
of the genus Macropus or Kangaroos, and of a powerful creature called
Thylacoleo, or Pouched Lion, which must be admitted as a new section of
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