long line of treacherous honey-glands by which the sarracenia entices
flies into the festering manure-wells of its sticky pitchers. The
minuteness and skill of all his observations on these lesser problems of
natural selection inevitably inspired faith among outsiders in the
cautious judgment of the observer and experimenter; and day by day
throughout his later years the evidence of the popular acceptance of his
doctrine, and of the dying away of the general ridicule with which it
was first received by the unlearned public, was very gratifying to the
great naturalist.
A year later, in 1876, came the 'Effects of Cross and Self Fertilisation
in the Vegetable Kingdom.' So far as regarded the world of plants,
especially with respect to its higher divisions, this work was of
immense theoretical importance; and it also cast a wonderful side-light
upon the nature of that strange distinction of sex which occurs both in
the vegetable and animal kingdom, and in each is the concomitant--one
might almost say the necessary concomitant--of high development and
complex organisation. The great result attained by Darwin in his long
and toilsome series of experiments on this interesting subject was the
splendid proof of the law that cross-fertilisation produces finer and
healthier offspring, while continuous self-fertilisation tends in the
long run to degradation, degeneration, and final extinction.
Here as elsewhere, however, Darwin's principle does not spring
spontaneous, like Athene from the head of Zeus, a goddess full-formed,
uncaused, inexplicable: it arises gradually by a slow process of
development and modification from the previous investigations of earlier
biologists. At the close of the last century, in the terrible year of
upheaval 1793, a quiet German botanist, Christian Konrad Sprengel by
name, published at Berlin his long unheeded but intensely interesting
work on the 'Fertilisation of Flowers.' In the summer of 1789, while all
Europe was ablaze with the news that the Bastille had been stormed, and
a new era of humanity begun, the calm and peaceful Pomeranian observer
was noting in his own garden the curious fact that many flowers are
incapable of being fertilised without the assistance of flying insects,
which carry pollen from the stamens of one blossom to the sensitive
surface or ovary of the next. Hence he concluded that the secretion of
honey or nectar in flowers, the contrivances by which it is protected
from rain, the bright hues or lines of the corolla, and the sweet
perfume distilled by the blossoms, are all so many cunning devices of
nature to ensure fertilisation by the insect-visitors. Moreover,
Sprengel observed that many flowers are of one sex only, and that in
several others the sexes do not mature simultaneously; 'so that,' said
he, 'nature seems to intend that no flower shall be fertilised by means
of its own pollen.' Indeed, in some instances, as he showed by
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