Animal behavior; Animals -- Color; Plants -- Color
The wing area may be divided into the strong costal margin, the hind
margin, the nervules, and the spaces; and, however complex the pattern
may be, it is always based upon these structure lines.
In the majority of insects the costal margin is marked with strong
colour. This may be noticed in _Papilio Machaon_, _P. merope_, _Vanessa
antiopa_, and the whites in Plate IV. The extreme tip of the fore-wings
is nearly always marked with colour, though this may run into the border
pattern. This colour is dark or vividly bright, and we know no
butterfly, not even dark ones, that has a light tip to the wings.
Sometimes, it is true, the light bead-border spots run to the tip, but
these are not cases in point. The development of tips has been traced in
Chapter VI., and need not be repeated.
The hind margin of both wings is very commonly emphasized by a border,
of which _V. Antiopa_, Pl. III. Fig. 3, is a very perfect example.
The border pattern may consist of one or more rows of spots, lines,
bands, or scallops;[31] and there is frequently a fine fringe, which in
many cases is white, with black marks, and to which the term
bead-pattern may be applied.
A definite relation subsists in most cases between the shape of the hind
margin and the character of the border-pattern. The plain or simple
bordered wings have plain border patterns, and the scalloped wings have
scalloped borders; or rather scalloped borders are almost exclusively
confined to scalloped wings. In our English butterflies, for instance,
out of the 62 species:--
33 have plain margins to the wings. In all the border is plain, or
wanting.
20 have the fore-wings plain, and the hind-wings scalloped, and in
all the hind-wings are scalloped and the fore-wings plain, or with
slightly scalloped border-patterns.
9 have scalloped margins and scalloped border-patterns.
Another relation between structure and pattern is found in those insects
which have tailed hind-wings, for the tail is very frequently emphasized
by a spot, often of a different colour from the rest of the wing as in
the Swallow-Tails, Plates IV. and V.
Yet another point may be noticed. In each wing there is a space, the
discoidal cell, _j_ Fig. 3, at the apex of which several nervures join,
forming knots. These are points at which obstacles exist to the flow of
the contents, and they are almost always marked by a distinct pattern.
We thus have a discoidal spot in very many butterflies, in nearly all
moths; and in the other orders of winged insects the decoration is even
more pronounced, as any one may see who looks at our dragon-flies,
wasps, bees, or even beetles.
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