Contributions to the Theory of Natural Selection: A Series of EssaysWallace, Alfred Russel
Science
Contributions to the Theory of Natural Selection: A Series of Essays
Wallace, Alfred Russel
Natural selection
The phenomena of dimorphism and polymorphism may be well illustrated by
supposing that a blue-eyed, flaxen-haired Saxon man had two wives, one a
black-haired, red-skinned Indian squaw, the other a woolly-headed,
sooty-skinned negress--and that instead of the children being mulattoes
of brown or dusky tints, mingling the separate characteristics of their
parents in varying degrees, all the boys should be pure Saxon boys like
their father, while the girls should altogether resemble their mothers.
This would be thought a sufficiently wonderful fact; yet the phenomena
here brought forward as existing in the insect-world are still more
extraordinary; for each mother is capable not only of producing male
offspring like the father, and female like herself, but also of
producing other females exactly like her fellow-wife, and altogether
differing from herself. If an island could be stocked with a colony of
human beings having similar physiological idiosyncrasies with Papilio
Pammon or Papilio Ormenus, we should see white men living with yellow,
red, and black women, and their offspring always reproducing the same
types; so that at the end of many generations the men would remain pure
white, and the women of the same well-marked races as at the
commencement.
The distinctive character therefore of dimorphism is this, that the
union of these distinct forms does not produce intermediate varieties,
but reproduces the distinct forms unchanged. In simple varieties, on the
other hand, as well as when distinct local forms or distinct species are
crossed, the offspring never resembles either parent exactly, but is
more or less intermediate between them. Dimorphism is thus seen to be a
specialized result of variation, by which new physiological phenomena
have been developed; the two should therefore, whenever possible, be
kept separate.
3. _Local form, or variety._--This is the first step in the transition
from variety to species. It occurs in species of wide range, when groups
of individuals have become partially isolated in several points of its
area of distribution, in each of which a characteristic form has become
more or less completely segregated. Such forms are very common in all
parts of the world, and have often been classed by one author as
varieties, by another as species. I restrict the term to those cases
where the difference of the forms is very slight, or where the
segregation is more or less imperfect. The best example in the present
group is Papilio Agamemnon, a species which ranges over the greater part
of tropical Asia, the whole of the Malay archipelago, and a portion of
the Australian and Pacific regions. The modifications are principally of
size and form, and, though slight, are tolerably constant in each
locality. The steps, however, are so numerous and gradual that it would
be impossible to define many of them, though the extreme forms are
sufficiently distinct. Papilio Sarpedon presents somewhat similar but
less numerous variations.
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