Contributions to the Theory of Natural Selection: A Series of EssaysWallace, Alfred Russel
Science
Contributions to the Theory of Natural Selection: A Series of Essays
Wallace, Alfred Russel
Natural selection
In Dr. Carpenter's
"Introduction to the Study of the Foraminifera," he states that "_there
is not a single specimen of plant or animal of which the range of
variation has been studied by the collocation and comparison of so large
a number of specimens as have passed under the review of Messrs.
Williamson, Parker, Rupert Jones, and myself, in our studies of the
types of this group_;" and the result of this extended comparison of
specimens is stated to be, "_The range of variation is so great among
the Foraminifera as to include not merely those differential characters
which have been usually accounted_ SPECIFIC, _but also those upon which
the greater part of the_ GENERA _of this group have been founded, and
even in some instances those of its_ ORDERS" (Foraminifera, Preface, x).
Yet this same group had been divided by D'Orbigny and other authors into
a number of clearly defined _families_, _genera_, and _species_, which
these careful and conscientious researches have shown to have been
almost all founded on incomplete knowledge.
Professor DeCandolle has recently given the results of an extensive
review of the species of Cupuliferae. He finds that the best-known
species of oaks are those which produce most varieties and subvarieties;
that they are often surrounded by provisional species; and, with the
fullest materials at his command, two-thirds of the species he considers
more or less doubtful. His general conclusion is, that "_in botany the
lowest series of groups,_ SUBVARIETIES, VARIETIES, _and_ RACES _are very
badly limited; these can be grouped into_ SPECIES _a little less vaguely
limited, which again can be formed into sufficiently precise_ GENERA."
This general conclusion is entirely objected to by the writer of the
article in the "Natural History Review," who, however, does not deny its
applicability to the particular order under discussion, while this very
difference of opinion is another proof that difficulties in the
determination of species do not, any more than in the higher groups,
vanish with increasing materials and more accurate research.
Another striking example of the same kind is seen in the genera Rubus
and Rosa, adduced by Mr. Darwin himself; for though the amplest
materials exist for a knowledge of these groups, and the most careful
research has been bestowed upon them, yet the various species have not
thereby been accurately limited and defined so as to satisfy the
majority of botanists. In Mr. Baker's revision of the British Roses,
just published by the Linnaean Society, the author includes under the
single species Rosa canina, no less than twenty-eight named _varieties_,
distinguished by more or less constant characters and often confined to
special localities; and to these are referred about seventy of the
_species_ of Continental and British botanists.
Public-domain text, read in full here on John Shaqi.
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