Contributions to the Theory of Natural Selection: A Series of EssaysWallace, Alfred Russel
Science
Contributions to the Theory of Natural Selection: A Series of Essays
Wallace, Alfred Russel
Natural selection
In estimating these numbers I have had the usual difficulty to
encounter, of determining what to consider species and what varieties.
The Malayan region, consisting of a large number of islands of generally
great antiquity, possesses, compared to its actual area, a great number
of distinct forms, often indeed distinguished by very slight
characters, but in most cases so constant in large series of specimens,
and so easily separable from each other, that I know not on what
principle we can refuse to give them the name and rank of species. One
of the best and most orthodox definitions is that of Pritchard, the
great ethnologist, who says, that "_separate origin and distinctness of
race, evinced by a constant transmission of some characteristic
peculiarity of organization_," constitutes a species. Now leaving out
the question of "origin," which we cannot determine, and taking only the
proof of separate origin, "_the constant transmission of some
characteristic peculiarity of organization_," we have a definition which
will compel us to neglect altogether the _amount_ of difference between
any two forms, and to consider only whether the differences that present
themselves are _permanent_. The rule, therefore, I have endeavoured to
adopt is, that when the difference between two forms inhabiting separate
areas seems quite constant, when it can be defined in words, and when it
is not confined to a single peculiarity only, I have considered such
forms to be species. When, however, the individuals of each locality
vary among themselves, so as to cause the distinctions between the two
forms to become inconsiderable and indefinite, or where the differences,
though constant, are confined to one particular only, such as size,
tint, or a single point of difference in marking or in outline, I class
one of the forms as a variety of the other.
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